Imperial Shags (Phalacrocorax (atriceps) atriceps) at breeding colony on small islets off the city of Ushuaia, Argentina, South America

imperial shags phalacrocorax atriceps breeding colony small islets ushuaia argentina south america nesting season wildlife seabirds marine life colonies nature birdwatching habitat mating eggs chicks environment natural behavior migration coastal biodiversity ecosystems avian shorebirds animal behavior reproduction plumage social behavior aquatic group dynamics adaptation argentine region habitat conservation flight foraging feeding nesting sites predator avoidance species interaction argentina wildlife breeding grounds conservation efforts ecological balance animal species ecological studies