Adult male mudskipper (Periophthalmus spp,) territorial display at low tide, Bako National Park, Sarawak, Borneo, Malaysia, Southeast Asia, Asia

photography colour colour image color color image horizontal horizontal image outdoors outside day nobody no one no one travel destination travel destinations travel adult animal amphibious male animal mudskipper periophthalmus territorial display bako national park national park sarawak borneo malaysia malaysian southeast asia south east asia southeast asian south east asian asia asian animal themes animal animals fauna one animal animal theme fish animal portrait wild animal wild animals wild animals in the wild wildlife wild life wild life focus on foreground mud texture fin spine spinal side view spotted amphibian