Female mantled howler monkey (Alouatta palliata) eating tree leaves in a north Pacific coast forest, Esperanza, Nosara, Guanacaste, Costa Rica

alouatta palliata tree leaves foraging costa rica guanacaste female monkey rainforest wildlife primate nature eating lush foliage animal behavior tropical forest herbivore green canopy biodiversity habitat social diet conservation ecological balance natural environment mammals forested area observation feeding jungle endangered species hotspot central america ecology ecosystem exploration leaf eater kingdom