Spotted sandpiper (Actitis macularius) non-breeding plumage, common shorebird in the Americas, Nosara River, Nosara, Guanacaste, Costa Rica

spotted sandpiper shorebird actitis macularius riverbank nosara river guanacaste costa rica wildlife foraging autumn nature birdwatching aquatic habitat food searching migratory bird natural environment brown plumage coastal small wetlands rocky shore avian behavior riverside insect prey ecological balance biodiversity observation landscape waterfowl animal ecosystem fauna conservation riverine tropical ornithology outdoor