Atlantic Puffin (Fratercula arctica) displaying, Skomer island, Pembrokeshire, Wales

fratercula arctica seaside sky sunset adult couple posing shout courtship behaviour profile shot overview backlighting piscivore temperate sea evening one on one fratercula atlantic puffin (fratercula arctica) vulnerable (iucn) vu national park may north atlantic ocean pembrokeshire coast national park call nobody coasts seasides coast skies sunsets adults posed pose settlings settling shouts shouting exclaims exclaim exclaiming parading parade parades seducing courtship behaviours seduce attractive seduction profile sights profile sight profile shots profile view profile views side view overviews global sight global sights full length backlightings backlit ichtyophage piscivores ichtyophagous temperate seas temperate ocean temperate oceans day fall nightfall late afternoon face to face atlantic puffins (fratercula arctica) vulnerable species national parks sensitive natural areas northern atlantic north atlantic calls with nobody strand area meteorological phenomenon meteorology science sun space stage of development individual pair enumeration vocal communication sound communication communicate social behaviour behavior reproductive behaviour shot light effect diet pattern species sea marine area twilight ambiance alcidae charadriiformes aves sauropsida tetrapoda tetrapodomorpha rhipidistia sarcopterygii euteleostomi gnathostomata vertebrata chordata deuterostomia coelomata bilateria metazoa opisthokonta eukaryota puffin (macareux) sea bird bird sauropsid vertebrate chordate wild animal animal threatened endangered species iucn red list of threatened species environment protection environment management biodiversity management environment atlantic ocean world sea and ocean pembrokeshire county wales (country) great britain united kingdom european union europe