Atlantic Puffin (Fratercula arctica) displaying, Skomer island, Pembrokeshire, Wales

fratercula arctica seaside sky sunset adult couple flap wings posing courtship behaviour overview backlighting piscivore temperate sea three quarter shot evening fratercula atlantic puffin (fratercula arctica) vulnerable (iucn) vu national park may north atlantic ocean pembrokeshire coast national park nobody coasts seasides coast skies sunsets adults flapping wings flaps wings flutter posed pose settlings settling parading parade parades seducing courtship behaviours seduce attractive seduction overviews global sight global sights full length backlightings backlit ichtyophage piscivores ichtyophagous temperate seas temperate ocean temperate oceans three quarter views three quarter view three quarter shots three quarter sight three quarter sights three quarter length day fall nightfall late afternoon atlantic puffins (fratercula arctica) vulnerable species national parks sensitive natural areas northern atlantic north atlantic with nobody strand area meteorological phenomenon meteorology science sun space stage of development individual pair enumeration locomotion motion reproductive behaviour behavior shot light effect diet pattern species sea marine area twilight ambiance alcidae charadriiformes aves sauropsida tetrapoda tetrapodomorpha rhipidistia sarcopterygii euteleostomi gnathostomata vertebrata chordata deuterostomia coelomata bilateria metazoa opisthokonta eukaryota puffin (macareux) sea bird bird sauropsid vertebrate chordate wild animal animal threatened endangered species iucn red list of threatened species environment protection environment management biodiversity management environment atlantic ocean world sea and ocean pembrokeshire county wales (country) great britain united kingdom european union europe