Mantled howler (Alouatta palliata) adult male howling on his territory, Pacific coast, Costa Rica

alouatta palliata tropical rainforest adult male alone branch foliage overview costa rica alouatta three quarter shot shrill cites appendix 1 least concern (iucn) lc mantled howler monkey (alouatta palliata) nobody tropical rain forests ombrophile forest shade plants rain forest rainforest rain forests ombrophile forests adults males single lonely only solo one lone boughts branches bought foliages overviews global sight global sights full length three quarter views three quarter view three quarter shots three quarter sight three quarter sights three quarter length holler holling shrilling least concern species with nobody tropical forest forest area stage of development individual enumeration morphology (botany) morphology organism leaf shot central america america atelidae platyrrhini haplorhini primates mammalia cynodontia therapsida eupelycosauria synapsida tetrapoda tetrapodomorpha rhipidistia sarcopterygii euteleostomi gnathostomata vertebrata chordata deuterostomia coelomata bilateria metazoa opisthokonta eukaryota shout vocal communication sound communication communicate social behaviour behavior convention on international trade in endangered species of wild species low risk (iucn) lr iucn red list of threatened species howler monkey new world monkey primate mammal therapsid synapsid vertebrate chordate wild animal animal