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Pair of Patagonian crested ducks (Lophonetta specularioides) in courtship behaviour, the Neck, Saunders Island, Falkland Islands, South America

Rockhopper penguin (Eudyptes chrysocome) mutual preening behaviour, Rockhopper Point, Sea Lion Island, Falkland Islands, South America

Great Blue Herons (Ardea herodias), the largest North American heron, standing in the nest, United States of America, North America

Dominant male proboscis monkey (Nasalis larvatus) on the lookout for challenges from younger males in the bachelor group, Labuk Bay Proboscis Monkey Sanctuary, Sabah, Borneo, Malaysia, Southeast Asia, Asia

Dominant male proboscis monkey (Nasalis larvatus) in aggressive display to other males, Labuk Bay Proboscis Monkey Sanctuary, Sabah, Borneo, Malaysia, Southeast Asia, Asia

Lion (Panthera leo) scratching and marking territory, Serengeti National Park, Tanzania, East Africa, Africa

Courting black-browed albatross (black-browed mollymawk) (Diomedea melanophris), West Point, Falkland Island, South America

Male Magnificent Frigatebird (Fregata magnificens) with inflated gular sac, North Seymour Island, Galapagos National Park, Ecuador, South America

Two male Cape giraffe (Giraffa camelopardalis giraffa) fighting, Imfolozi Game Reserve, South Africa, Africa

Two African elephant (Loxodonta africana) face to face, Addo Elephant National Park, South Africa, Africa

Male impala (Aepyceros melampus) with a red-billed oxpecker (Buphagus erythrorhynchus), Kruger National Park, South Africa, Africa

Wild burro (donkey) (Equus asinus) (Equus africanus asinus) jenny biting its foal's ear, Custer State Park, South Dakota, United States of America, North America

Pink salmon (Oncorhynchus gorbuscha), spawning season, Port Valdez, Alaska, United States of America, North America

Baby cleaning a male Gelada baboon, Simien Mountains National Park, UNESCO World Heritage Site, Ethiopia, Africa

Olive Baboon (Papio cynocephalus anubis) juvenile grooming an adult male, Ngorongoro Crater, Tanzania, East Africa, Africa

Wattled Starling (Creatophora cinerea), male in breeding plumage, Kruger National Park, South Africa, Africa

Portrait of male Great Frigatebird (Fregata minor) on Genovesa Island, Galapagos National Park, Ecuador, South America

Bull moose (Alces alces) calling, Wasatch Mountain State Park, Utah, United States of America, North America

Impala (Aepyceros melampus) buck with red-billed oxpecker (Buphagus erythrorhynchus), Kruger National Park, South Africa, Africa

Adult Chilean skua, Stercorarius chilensis, harassing kelp gulls to force regurgitation, Beagle Channel, Argentina, South America

Ground squirrels (Geosciurus inauris), Kgalagadi Transfrontier Park, Northern Cape, South Africa, Africa

Adult Chilean skua, Stercorarius chilensis, harassing kelp gulls to force regurgitation, Beagle Channel, Argentina, South America

Black-browed albatross, Thalassarche melanophris, courtship display on New Island, Falkland Islands, South Atlantic Ocean

Portrait of Nazca Boobies (Sula granti), Genovesa Island, Galapagos National Park, Ecuador, South America

Olive baboon (Papio cynocephalus anubis) juvenile grooming an adult male, Ngorongoro Crater, Tanzania, East Africa, Africa

New Zealand Sea Lions (Phocarctos hookeri) at Otago Peninsula, Dunedin, Otago, South Island, New Zealand, Pacific

Male Magnificent Frigatebird (Fregata magnificens) with inflated gular sac, North Seymour Island, Galapagos National Park, Ecuador, South America

South American sea lion bull, Otaria flavescens, attempting to mate on a small islet in the Beagle Channel, Argentina, South America

Small American black bear (Ursus americanus) with cinnamon or blond colouring, digging tree bark for insects, Crescent Meadow trail, Sequoia National Park, Sierra Nevada, California, United States of America, North America

Yellow pine chipmunk (Eutamias amoenus) eating sagebrush leaves, Yellowstone National Park, Wyoming, United States of America, North America

Bison (Bison bison) bull demonstrating the flehmen response, Yellowstone National Park, Wyoming, United States of America, North America

A pair of red phalaropes (Phalaropus fulicarius) in courtship display at Russebuhkta, Edgeoya, Svalbard, Arctic, Norway, Europe

Rockhopper penguin (Eudyptes chrysocome) courtship behaviour, Rockhopper Point, Sea Lion Island, Falkland Islands, South America

Portrait of male Great Frigatebird (Fregata minor) on Genovesa Island, Galapagos National Park, Ecuador, South America

Black-browed albatross, Thalassarche melanophris, courtship display on New Island, Falkland Islands, South Atlantic Ocean

Great Blue Heron (Ardea herodias), the largest North American heron, in breeding display, United States of America, North America

A raft of sea otters, Enhydra lutris, grooming their fur in kelp in the Inian Islands, Southeast Alaska, United States of America

Male Magnificent Frigatebird (Fregata magnificens) with inflated gular sac, North Seymour Island, Galapagos National Park, Ecuador, South America

Male impala (Aepyceros melampus) with a red-billed oxpecker (Buphagus erythrorhynchus), Kruger National Park, South Africa, Africa

Black-browed albatross, Thalassarche melanophris, courtship display on New Island, Falkland Islands, South Atlantic Ocean

Male Magnificent Frigatebird (Fregata magnificens) with inflated gular sac, North Seymour Island, Galapagos National Park, Ecuador, South America

A rafting sea otter, Enhydra lutris, grooming its fur in kelp in the Inian Islands, Southeast Alaska, United States of America

A rafting sea otter, Enhydra lutris, grooming its fur in kelp in the Inian Islands, Southeast Alaska, United States of America

Cape mountain zebra (Equus zebra zebra) sparring, Mountain Zebra National Park, South Africa, Africa

Immature red-billed oxpecker (Buphagus erythrorhynchus) on an impala, Kruger National Park, South Africa, Africa

Two blesbok (Damaliscus pygargus phillipsi) facing off, Mountain Zebra National Park, South Africa, Africa

Adult razorbills (Alca torda) at the abandoned fishing settlement at Mastad on the island of Voroya, Norway, Scandinavia, Europe

Adult pink salmon, Oncorhynchus gorbuscha, returning to the Indian River to spawn near Sitka, Alaska, United States of America

View of a male pink salmon, Oncorhynchus gorbuscha, in the Indian River spawning near Sitka, Alaska, United States of America

Two teenaged male African elephant (Loxodonta africana) playing, Kruger National Park, South Africa, Africa

Cave Myotis (Myotis velifer) in flight in captivity, Hidalgo County, New Mexico, United States of America, North America

Male impala (Aepyceros melampus) with two red-billed oxpeckers (Buphagus erythrorhynchus), Kruger National Park, South Africa, Africa

Desert elephants (Loxodonta africana), Huab River Valley, Torra Conservancy, Damaraland, Namibia, Africa

California sea lions (Zalophus californianus), thermoregulating, Isla San Marcos, Baja California Sur, Mexico, North America

Young African elephant (Loxodonta africana) playing in the water, Serengeti National Park, UNESCO World Heritage Site, Tanzania, East Africa, Africa

Desert elephant (Loxodonta africana), Huab River Valley, Torra Conservancy, Damaraland, Namibia, Africa

Female bearded woodpecker (Dendropicos namaquus), Serengeti National Park, Tanzania, East Africa, Africa

Two young African elephant (Loxodonta africana) playing in the water, Serengeti National Park, UNESCO World Heritage Site, Tanzania, East Africa, Africa

Two bull elk (Cervus canadensis) sparring during the rut, Jasper National Park, UNESCO World Heritage Site, Alberta, Canada, North America

Common zebra (Burchell's zebra) (Equus burchelli) dust bathing, Ngorongoro Crater, Tanzania, East Africa, Africa

Desert elephants (Loxodonta africana), Huab River Valley, Torra Conservancy, Damaraland, Namibia, Africa

Baby elephant in a cloud of dust sprayed by its mother, Amboseli National Park, Kenya, East Africa, Africa

Juvenile grey heron (Ardea cinerea) in flight, escaping from an aggressive rival in a territorial dispute, Rutland Water, Rutland, England, United Kingdom, Europe

Chacma baboon (Papio cynocephalus ursinus), eating fruit of Sausage Tree (Kigelia africana), Kruger National Park, South Africa, Africa

Adult brown bears (Ursus arctos) courtship behaviour, Brooks River, Katmai National Park near Bristol Bay, Alaska, United States of America

Agonistic behaviour in a tight group of moulting southern elephant seals (Mirounga leonina), Grytviken, South Georgia, Polar Regions

Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa, Africa

Blue-footed booby (Sula nebouxii) pair in courtship display on San Cristobal Island, Galapagos, Ecuador, South America

Blue-footed booby (Sula nebouxii) pair in courtship display on San Cristobal Island, Galapagos, Ecuador, South America

Blue-footed booby (Sula nebouxii) pair in courtship display on North Seymour Island, Galapagos, Ecuador, South America

Adult male great frigatebirds (Fregata minor), courtship display. Genovesa Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Adult Sally lightfoot crab (Grapsus grapsus) preparing to molt on Fernandina Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild horses playing and grazing and Khangai mountains in the background, Hovsgol province, Mongolia, Central Asia, Asia