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Puffin (Fratercula arctica) in flight during high winds with ruffled feathers, Farne Islands, Northumberland, England

Puffin (Fratercula arctica) on a rocky outcrop on the Farne Islands, Northumberland, England, United Kingdom, Europe

Puffin (Fratercula arctica) at Latrabjarg, largest bird colony in Europe, West Fjords (Vestfirdir), Iceland, Polar Regions

Adult Atlantic puffin (Fratercula arctica), with small fish caught in its bill, Grimsey Island, Iceland, Polar Regions

Adult Atlantic puffin (Fratercula arctica), with small fish caught in its bill, Grimsey Island, Iceland, Polar Regions

Puffins (Fratercula arctica) on small cliffs near Bakkagerdi (Borgafjordur Eystri), East Fjords (Austurland), Iceland, Polar Regions

Puffin (Fratercula arctica) at rest with a beak full of sand eels, Farne Islands, Northumberland, England, United Kingdom, Europe

Puffin (Fratercula arctica), Heimaey Island, Vestmannaeyjar, volcanic Westman Islands, Iceland, Polar Regions

Puffin (Fratercula arctica) walking to its burrow along a rock with a beak full of sand eels, Farne Islands, Northumberland, England, United Kingdom, Europe

Adult Atlantic puffin (Fratercula arctica) with small fish, Vigur Island, off the west coast of Iceland, Polar Regions

Puffins (Fratercula arctica) at Latrabjarg, largest bird colony in Europe, West Fjords (Vestfirdir), Iceland, Polar Regions

Adult Atlantic puffins (Fratercula arctica) on Vigur Island, off the west coast of Iceland, Polar Regions

Puffin (Fratercula arctica) hanging onto the rock on a summer's evening on the Saltees Islands, County Wexford, Leinster, Republic of Ireland, Europe

Puffin (Fratercula arctica), Heimaey Island, Vestmannaeyjar, volcanic Westman Islands, Iceland, Polar Regions

Atlantic puffins (Fratercula arctica), Noss Island, Shetland Islands, Scotland, United Kingdom, Europe

Puffin (Fratercula arctica) with sandeels, Farne Islands, Northumberland, England, United Kingdom, Europe

Juvenile puffin (Fratercula arctica) on the water near the island of Bolscheoya in the Svalbard Archipelago, Norway, Arctic, Europe

A pair of Atlantic Puffins (Fratercula arctica), on a hilltop in Borgarfjaroarhofn, Iceland, Polar Regions

Black and white image of an Atlantic Puffin (Fratercula arctica), at sunset in Borgarfjaroarhofn, Iceland, Polar Regions

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Atlantic Puffin (Fratercula arctica) in its burrow on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Adult Atlantic puffins (Fratercula arctica), breeding on the island of Bolscheoya in the Svalbard Archipelago, Norway

Puffin (Fratercula arctica) with sand eels, Farne Islands, Northumberland, England, United Kingdom, Europe

Atlantic puffin (Fratercula arctica), Sumburgh Head, South Mainland, Shetland Islands, Scotland, United Kingdom, Europe

Atlantic puffin (Fratercula arctica), Sumburgh Head, South Mainland, Shetland Islands, Scotland, United Kingdom, Europe

Black and whilte image of an Atlantic Puffin (Fratercula arctica), after having jumped from a cliff edge in Borgarfjaroarhofn, Iceland, Polar Regions

An Atlantic Puffin (Fratercula arctica), with twigs in its beak in Borgarfjaroarhofn, Iceland, Polar Regions

Black and white image of an Atlantic Puffin (Fratercula arctica), in Borgarfjaroarhofn, Iceland, Polar Regions

Black and white image of an Atlantic Puffin (Fratercula arctica), at sunset in Borgarfjaroarhofn, Iceland, Polar Regions

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Adult Atlantic puffins (Fratercula arctica), breeding on the island of Bolscheoya in the Svalbard Archipelago, Norway

Adult Atlantin puffin (Fratercula arctica) during breeding season, Isfjardardjup, Westfjords region, Northwestern Iceland

Adult Atlantin puffin (Fratercula arctica) during breeding season, Isfjardardjup, Westfjords region, Northwestern Iceland

Adult Atlantin puffin (Fratercula arctica) during breeding season, Isfjardardjup, Westfjords region, Northwestern Iceland

Adult Atlantin puffin (Fratercula arctica) during breeding season, Isfjardardjup, Westfjords region, Northwestern Iceland

Adult Atlantin puffin (Fratercula arctica) during breeding season on Sumburgh Head, Shetlands, Scotland, United Kingdom

Adult Atlantin puffin (Fratercula arctica) during breeding season on Sumburgh Head, Shetlands, Scotland, United Kingdom

Adult Atlantin puffin (Fratercula arctica) during breeding season, Isfjardardjup, Westfjords region, Northwestern Iceland

Adult Atlantin puffin (Fratercula arctica) during breeding season on Sumburgh Head, Shetlands, Scotland, United Kingdom

Black and white image of an Atlantic Puffin, Fratercula arctica, in Borgarfjaroarhofn, Iceland, Polar Regions

An Atlantic Puffin (Fratercula arctica), landing on a cliff in Borgarfjaroarhofn, Iceland, Polar Regions

An Atlantic Puffin (Fratercula arctica), on a cliff edge in Borgarfjaroarhofn, Iceland, Polar Regions

An Atlantic Puffin (Fratercula arctica), atop a cliff with an Iceland gull (Larus glaucoides), soaring above in Westman Islands, Iceland, Polar Regions

A pair of Atlantic Puffins (Fratercula arctica), on a cliff edge in Westman Islands, Iceland, Polar Regions

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Atlantic Puffin (Fratercula arctica), on Skomer Island in July, a nature reserve off the coast of Pembrokeshire, Wales, United Kingdom, Europe

Puffin (Fratercula arctica) with sand eels, Farne Islands, Northumberland, England, United Kingdom, Europe

Atlantic puffin (Fratercula arctica), taking flight on Bjornoya, Svalbard, Norway,.Scandinavia, Europe

Puffin (Fratercula arctica) with sand eels, Farne Islands, Northumberland, England, United Kingdom, Europe

Adult Atlantic puffin (Fratercula arctica), returning to the nest site with fish (sand eels) for its chick on Grimsey Island, Iceland, Polar Regions

Adult Atlantic puffin (Fratercula arctica), returning to the nest site with fish (sand eels) for its chick on Grimsey Island, Iceland, Polar Regions

Adult Atlantic puffin (Fratercula arctica) at Sumburgh Head, Mainland Island, Shetland Isles, Scotland, United Kingdom, Europe

Atlantic puffin (Fratercula arctica), Sumburgh Head, South Mainland, Shetland Islands, Scotland, United Kingdom, Europe

Adult Atlantic puffin (Fratercula arctica), returning to the nest site with fish (sand eels) for its chick on Grimsey Island, Iceland, Polar Regions

Atlantic puffins (Fratercula arctica) take flight from a cliff-top, Inner Farne, Farne Islands, Northumberland, England, United Kingdom, Europe

Adult Atlantic puffin (Fratercula arctica) in flight with fish in its bill, Snaefellsnes Peninsula, Iceland, Polar Regions

Puffin (Fratercula arctica) with sand eels, Farne Islands, Northumberland, England, United Kingdom, Europe