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Close-up portrait of a Bengal tiger (Panthera tigris tigris) standing in the forest looking up, Madhya Pradesh, India

Close-up portrait of a lioness (Panthera leo) lying down in the golden long grass on the savanna in the Etosha National Park; Otavi, Oshikoto, Namibia

A leopard, Panthera pardus, lies on a broken tree branch, drapes its feet and tail over the branch, looking away, ears back, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A pride of lion, Panthera leo, and their cubs walk through a river causeway, ears back, wound on one leg, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, turns over its right shoulder, looking out, ears back, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, walks towards the camera on sand road, direct gaze, ears facing backward, mouth open, long green grass, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, walks towards the camera on a sand road, looking away, ears back, mouth open, in black and white, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, lies on a dead tree branch, tail drapes down, yawning with ears back and eyes closed, in black and white, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

The back of a head of a lioness, Panthera leo, ears back, front paw resting on tree trunk, claws showing, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, stands in a tree over a carcass, alert, blood on nose and snout, ears up, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard mother, Panthera pardus, lies down, eyes closing, mouth open as a cub bites her ear with blue eyes, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, sits in tall dry yellow grass looking around, ears facing forward, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard's ear and eye, Panthera pardus, ripped ear with ticks on, tick on eyelid, greenery in the background, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, sits in tall dry brown grass and looks around alert, ears facing forward, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard's head and front leg, Panthera pardus, lying on a tree branch, alert, ears forward, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, crouching low, stalks through dry grass, ears back, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, crouches down to drink water, direct gaze, ears back, ripples in water, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, lying down resting its head on a branch, ears back, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, lies in the grass, direct gaze, ears up, greenery in foreground, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, lies on the branch of a tree, legs tangling over the branch, alert, ears forward, head resting on branch, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, claws the trunk of a tree, ears back, looking out of frame, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, walks across a clearing, tail curled in the air, in the foreground a male impala with horns faces the leopard with ears perked, Aepyceros melampus, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

The head of a lioness, Panthera leo, lying down, alert, ears forward, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, walks towards the camera on a sand road, looking away, ears forward, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A leopard, Panthera pardus, lies on the ground in the sunlight, alert, ears forward, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa

A lioness, Panthera leo, with no tail walks across a river, ears back, looking away, splashing, sunset in the background, Londolozi Game Reserve, Sabi Sands, Greater Kruger National Park, South Africa