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Captive Galapagos giant tortoise (Chelonoidis nigra) at the Charles Darwin Research Station on Santa Cruz Island, Galapagos Islands, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus), Charles Darwin Research Station, Galapagos Islands, UNESCO World Heritge Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoises (Chelonoidis spp), Charles Darwin Research Station, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoises (Chelonoidis spp), Charles Darwin Research Station, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Close-up of foot of captive Galapagos giant tortoise (Chelonoidis spp), Charles Darwin Research Station, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Giant Tortoise at Charles Darwin Research Station, Puerto Ayora, Santa Cruz (Indefatigable) Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Chelonoidis spp), Charles Darwin Research Station, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Chelonoidis spp), Charles Darwin Research Station, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoises (Chelonoidis spp), Charles Darwin Research Station, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Charles Darwin Research Station, Puerto Ayora, Santa Cruz (Indefatigable) Island, Galapagos, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) with photographer at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Group, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Group, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Group, Ecuador

Photographers with captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador