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Male black grouse (Lyrurus tetrix) fighting on snow covered field, Finland
Adult black-browed albatross (Thalassarche melanophrys), pair in courtship display at nesting site on New Island, Falklands, South America
Adult black-browed albatross (Thalassarche melanophrys), trio in courtship display at nesting site on New Island, Falklands, South America
Southern elephant seal (Mirounga leonina) mating behavior on South Georgia Island, Southern Ocean
Adult Grey-headed Albatross (Thalassarche chrysostoma) courtship display at Elsehul, South Georgia
Black-browed albatross (Thalassarche melanophrys) breeding colony on Carcass Island in the Falkland Islands
Black-browed albatross (Thalassarche melanophrys) breeding colony on New Island in the Falkland Island.
Adult brown bears (Ursus arctos) courtship behaviour, Brooks River, Katmai National Park near Bristol Bay, Alaska, United States of America
Adult Nazca booby (Sula grantii) courtship behavior in the Galapagos Island Archipelago, UNESCO World Heritage Site, Ecuador, South America
Blue Footed Booby (Sula nebouxii), a marine bird found in the Eastern Pacific whose unusual blue feet feature in courtship rituals, Galapagos, UNESCO World Heritage Site, Ecuador, South America
Southern elephant seal (Mirounga leonina) mating behavior on South Georgia Island, Southern Ocean
Adult arctic tern (Sterna paradisaea) male bringing female a fish, Spitsbergen Island in the Svalbard Archipelago, Norway
Imperial Shag (Phalacrocorax atriceps), pair exhibiting courtship behavior on New Island in the Falkland Islands, South America
Imperial Shag (Phalacrocorax atriceps), pair exhibiting courtship behavior on New Island in the Falkland Islands, South America
Male Great frigatebird (Fregata minor) in breeding plumage with red gular pouch, on North Seymour Island, Galapagos Islands, UNESCO World Heritage Site, Ecaudor, South America
Adult brown bear pair (Ursus arctos) exhibiting courtship behavior at Scidmore Cut in Glacier Bay National Park, Alaska, United States of America
Adult whooper swan (Cygnus cygnus), pair on nest near Lake Myvatn, Iceland
Aerial of cattle moving to a waterhole, Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Aerial of cattle moving to a waterhole, Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Aerial of cattle moving to a waterhole, Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Wodaabe-Bororo men with faces painted at the annual Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Wodaabe-Bororo men with faces painted at the annual Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Wodaabe-Bororo men with faces painted at the annual Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Wodaabe-Bororo men with faces painted at the annual Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Wodaabe-Bororo man with face painted at the annual Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Young girls at the Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Tuaregs on their camels, Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Young men arriving for the Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Wodaabe-Bororo man with face painted at the annual Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Local women arriving on the Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Wodaabe-Bororo men with faces painted at the annual Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Young girls at the Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Tuaregs on camels, Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Young girls selling food, Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Young girls at the Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
Colourful dressed girl at the Gerewol festival, courtship ritual competition among the Wodaabe Fula people, Niger, West Africa, Africa
A pair of Black guillemots bonding in Oban, Scotland, United Kingdom, Europe
Blue-footed booby (Sula nebouxii) pair in courtship ritual in the Galapagos Island Group, Ecuador. The Galapagos are a nesting and breeding area for blue-footed boobies.
Black-browed Albatross (Thalassarche melanophrys) mating ritual in the Falkland Islands.
Black Guillemot (Cepphus grylle) circling in a paring/mating ritual in the middle of Oban Bay with boats in the background. Black Guillemots nest in drains and holes in the sea wall in the middle of Oban town centre. Argyll, Scotland, UK
Green Sea Turtle (Chelonia mydas) mating ritual, Kaneohe Bay, Oahu, Hawaii, United States of America, Pacific
Green Sea Turtle (Chelonia mydas) mating ritual, Kaneohe Bay, Oahu, Hawaii, United States of America, Pacific
Adult California Gray Whale (Eschrichtius robustus) fluke-up dive - possible courtship behavior - in the calm waters of San Ignacio Lagoon, Baja, Mexico.
Cuttlefish (Sepia officinalis) Courtship. Babbacombe, Torquay, South Devon, UK. (A4 only).
Cuttlefish (Sepia officinalis) Two in courtship, being photographed by diver. Babbacombe, Torquay, South Devon, UK
(RR)
Close up of the bracelets, Gerewol festival, courtship ritual competition among the Woodaabe Fula people, Niger, Africa
Wodaabe-Bororo man with face painted at the annual Gerewol festival, courtship ritual competition among the Fulani ethnic group, Niger, Africa
Northern gannet (Morus bassanus), courtship and welcoming ritual on return of mate, Helgoland, Schleswig-Holstein, Germany, Europe
Young Peul man, Gerewol festival, courtship ritual competition among the Fulani ethnic group, Niger, Africa
Black-browed albatross or black-browed mollymawk (Thalassarche melanophris), typical courtship and greeting behaviour. South America, Falkland Islands, January
Black-browed albatross or black-browed mollymawk (Thalassarche melanophris), typical courtship and greeting behaviour. South America, Falkland Islands, January
A Pair Of Bald Eagles (Haliaeetus Leucocephalus) That Are Likely Mates Do A Bonding Ritual Of Flying Together And Touching Or Locking Talons Together And Then Tumbling Down Through The Air, South-Central Alaska, Alaska, United States Of America
Royal tern (Thalasseus maximus), Courtship ritual, Magdalena Bay (Madelaine Bay), Puerto San Carlos, Baja California Sur, Mexico