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Polar bear (Ursus maritimus) males playfighting, captive, Highland Wildlife Park, Kingussie, Scotland, United Kingdom, Europe

Juvenile grey heron (Ardea cinerea) in flight, escaping from an aggressive rival in a territorial dispute, Rutland Water, Rutland, England, United Kingdom, Europe

Adult bull southern elephant seals (Mirounga leonina) fighting for breeding grounds on South Georgia Island, Southern Ocean

Male and female (on right) Grey Seals (Halichoerus grypus) fighting on the shore, in winter, at Donna Nook Nature Reserve, on the North Sea coast of Lincolnshire, Great Britain

Adult brown bears (Ursus arctos) disputing fishing rights for salmon at the Brooks River, Katmai National Park, Alaska, United States of America

Mother polar bear (Ursus maritimus) and cub-of-the-year disputing then allowing a male bear to feed on a fin whale, Svalbard Archipelago, Norway

Adult brown bears (Ursus arctos) disputing fishing rights for salmon in the Brooks River, Katmai National Park, Alaska, United States of America

Adult Brunnich's guillemots (Uria lomvia) in a territorial dispute at Alkefjellet, Spitsbergen, Svalbard, Norway, Europe

A combative male iguana bloodied in territorial battle. Santiago Island, Galapagos.S(marine iguana)S(territorial dispute)S(Galapagos)S(UW)

Burchell's zebra (Equus burchelli) quarrelling at a waterhole, Etosha National Park, Namibia, Africa

Two young eagle (Haliaeetus albicilla), fighting in the air, with ravens (Corvus corax) at sead deer, Masuria, Poland, Europe

Two young eagles (Haliaeetus albicilla), fighting on the ground, with dead deer, Masuria, Poland, Europe

Hippos arguing (Hippopatamus amphibius) in the water, iSimangaliso Wetland Park, National Park, Kwazulu Natal, South Africa, Africa

Puffin (Fratercula arctica) in winter in snow, fighting, Hornoya Island, Hornoya, Vardo, Varanger Peninsula, Troms og Finnmark, Norway, Europe

On nest eating egg of Black-browed albatross (Thalassarche melanophris). Falkland Skua or Brown Skua (Stercorarius antarcticus, exact taxonomy is under dispute). They are the great skuas of the southern polar and subpolar region. South America, Falkland Islands, January

Chick. Falkland Skua or Brown Skua (Stercorarius antarcticus, exact taxonomy is under dispute). They are the great skuas of the southern polar and subpolar region. South America, Falkland Islands, January

Chick. Falkland Skua or Brown Skua (Stercorarius antarcticus, exact taxonomy is under dispute). They are the great skuas of the southern polar and subpolar region. South America, Falkland Islands, January

Chick. Falkland Skua or Brown Skua (Stercorarius antarcticus, exact taxonomy is under dispute). They are the great skuas of the southern polar and subpolar region. South America, Falkland Islands, January

Falkland Skua or Brown Skua (Stercorarius antarcticus, exact taxonomy is under dispute). They are the great skuas of the southern polar and subpolar region. South America, Falkland Islands, January

Falkland Skua or Brown Skua (Stercorarius antarcticus, exact taxonomy is under dispute). They are the great skuas of the southern polar and subpolar region. South America, Falkland Islands, January

Chick. Falkland Skua or Brown Skua (Stercorarius antarcticus, exact taxonomy is under dispute). They are the great skuas of the southern polar and subpolar region. South America, Falkland Islands, January

A pair of Pacific walruses (Odobenus rosmarus) joust with ivory tusks on the rocky shore of Walrus Islands State Game Sanctuary on Round Island at Bristol Bay in Alaska's Bering Sea.

Two threatening swallow tailed gulls, Creagrus furcatus, South america, Gal?pagos, Galapagos, Island

Gelada or Gelada baboon (Theropithecus gelada), fight between two males, Debre Libanos, Rift Valley, Ethiopia, Africa

Gelada or Gelada baboon (Theropithecus gelada), fight between two males, Debre Libanos, Rift Valley, Ethiopia, Africa

Southern Elephant Seal (Mirounga leonina). Elephant seals fighting on the beach of St Andrews Bay, South Georgia.

Hippopotamus (Hippopotamus amphibius) are playing to fight at dusk , Kenya , Masaï Mara, National Reserve

Middle Spotted Woodpecker (Dendrocopos medius) and Black bird (Turdus merula) fighting, Vosges du Nord Regional Natural Park, France

Two Giraffes doing necking parade in Kgalagadi transfrontier park, South Africa ; Specie Giraffa camelopardalis family of Giraffidae

Preying on Gentoo Penguins (Pygoscelis papua) hovering over colony.. Falkland Skua or Brown Skua (Stercorarius antarcticus, exact taxonomy is under dispute). They are the great skuas of the southern polar and subpolar region. South America, Falkland Islands, January

African elephant (Loxodonta africana) facing at Okaukuejo waterhole at sunset, Etosha National Park, Namibia

Falkland Skua or Brown Skua (Stercorarius antarcticus, exact taxonomy is under dispute). They are the great skuas of the southern polar and subpolar region. South America, Falkland Islands, January

Two White rhinoceros (Ceratotherium simum) are fighting with each other. Kenya. Nakuru National Park. Africa.

Two White rhinoceros (Ceratotherium simum) are fighting with each other. Kenya. Nakuru National Park. Africa.

Green Honeycreeper (Chlorophanes spiza) confronting a Saffron-crowned Tanager (Tangara xanthocephala), Colombia

Eastern moose (Alces americanus) males confront each other on a snowy forest track after the rutting season. A friendly fight that allows the dominant male to consolidate his stature. Parc de la Gaspesie. Quebec. Canada

Red fox (Vulpes vulpes) males in territorial confrontation. Montreal Botanical Garden. Quebec. Canada

Common Hippo (Hippopotamus amphibius), showdown, South Luangwa natioinal Park, Luangwa river, Zambia

Grevy's Zebra (Equus grevyi), in the savannah, Fight between males, dry shrubby savannah, Laïkipia County, Kenya, East Africa, Africa

Two giraffes (Giraffa camelopardalis tippelskirchi) are fighting each other in the savannah. Kenya. Tanzania. Eastern Africa.

Red Hartebeest (Alcelaphus buselaphus caama). Fighting bulls. Kalahari Desert, Kgalagadi Transfrontier Park, South Africa.

Springbok (Antidorcas marsupialis). Fighting males. Kalahari Desert, Kgalagadi Transfrontier Park, South Africa.

Red deer (Cervus elaphus) fighting on the island of Jura, Scotland, with sea and Islay in the background.

Two giraffes (Giraffa camelopardalis tippelskirchi) are fighting each other in the savannah. Kenya. Tanzania. Eastern Africa.

Muskox or Muskoxen (Ovibos moschatus) bulls fighting in deep snow during winter. Bardu, Polar Park, Norway, Scandinavia, Europe