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Galapagos white-cheeked pintail (Anas bahamensis galapagensis) feeding near giant tortoise in the Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoises (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoises (Chelonoidis spp), Charles Darwin Research Station, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoises (Chelonoidis spp), Charles Darwin Research Station, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoises (Chelonoidis spp), found in Rancho Manzanillo, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoises (Chelonoidis spp), Charles Darwin Research Station, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoises (Chelonoidis spp), found in Rancho Manzanillo, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) with photographer at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Young Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the tortuguero breeding station, Puerto Villamil on Isabela Island in the Galapagos Island Archipelago, Ecuador

Galapagos white-cheeked pintail (Anas bahamensis galapagensis) feeding near giant tortoise in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Photographers with captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) on the upslope grasslands of Cruz Island in the Galapagos Island Group, Ecuador

Galápagos tortoise or Galápagos giant tortoise (Geochelone nigra), adult feeding, Galapagos Islands, Pacific Ocean

Galápagos tortoise or Galápagos giant tortoise (Geochelone nigra), Galapagos Islands, Pacific Ocean

Galápagos tortoise or Galápagos giant tortoise (Geochelone nigra), adult, Galapagos Islands, Pacific Ocean

Portrait of Galapagos Giant Tortoise (Geochelone elephantopus) in the Charles Darwin Station in Puerto Ayora, Insel Santa Cruz, Galapagos Inseln, Galapagos Islands, Ecuador, South America

Galapagos giant tortoise (Chelonoidis nigra) lumbers slowly across a long, straight dirt road that stretches off to the horizon. Beyond the grass verge, there is dense forest on either side; Galapagos Islands, Ecuador

Galapagos giant tortoise (Chelonoidis nigra) lumbers slowly across a long, straight dirt road that stretches off to the horizon. Beyond the grass verge, there is dense forest on either side; Galapagos Islands, Ecuador