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Wild Galapagos tortoise (Geochelone elephantopus), Urbina Bay, Isabela Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos tortoise (Geochelone elephantopus), Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos tortoise (Geochelone elephantopus), Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Detail of carapace, wild Galapagos tortoise (Geochelone elephantopus), Urbina Bay, Isabela Island, Galapagos Islands, Ecuador, South America

Wild Galapagos tortoise (Geochelone elephantopus), Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos tortoise (Geochelone elephantopus), Urbina Bay, Isabela Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island, Galapagos Islands, UNESCO World Heritge Site, Ecuador, South America

Wild Galapagos tortoise (Geochelone elephantopus), Urbina Bay, Isabela Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos tortoise (Geochelone elephantopus), Urbina Bay, Isabela Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus), Charles Darwin Research Station, Galapagos Islands, UNESCO World Heritge Site, Ecuador, South America

Wild Galapagos tortoise (Geochelone elephantopus), Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Young Captive Galapagos giant tortoise (Geochelone elephantopus) being fed, San Cristobal Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) at Urbina Bay, Isabela Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) at Urbina Bay, Isabela Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) at Urbina Bay, Isabela Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) at Urbina Bay, Isabela Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) with cow on Santa Cruz Island, Galapagos, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoises (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus), face detail, Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Tourist photograhing a wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Tourists photographing a wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Tourist photographing a wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Tourist photographing a wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Tourist photographing a wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) in mud pit on Santa Cruz Island, Galapagos, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island, Galapagos, Ecuador, South America

Giant tortoise (Geochelone elephantopus vandenburghi), Isla Sant Cruz, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Giant tortoise (Geochelone elephantopus vandenburghi), Isla Sant Cruz, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Giant tortoise (Geochelone elephantopus vandenburghi), Isla Sant Cruz, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus), in Urbina Bay, Isabela Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus), with tourists on Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos tortoise (Geochelone elephantopus), Santa Cruz Island, Galapagos Islands, Ecuador, South America

Geochelone elephantopus vandenburghi, Giant Tortoise Breeding Centre, Puerto Villamil, Isla Isabela, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) with photographer at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) at Urbina Bay, Isabela Island in the Galapagos Island Archipelago, Ecuador

Young Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the tortuguero breeding station, Puerto Villamil on Isabela Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Group, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Group, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador