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Giant tortoise (Geochelone nigra) at the Galapaguera de Cerro Colorado, tortoise breeding center, Isla San Cristobal (San Cristobal Island), Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Giant tortoise (Geochelone nigra) at the Galapaguera de Cerro Colorado, tortoise breeding center, Isla San Cristobal (San Cristobal Island), Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) with photographer at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Young Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the tortuguero breeding station, Puerto Villamil on Isabela Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Photographers with captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Galápagos tortoise or Galápagos giant tortoise (Geochelone nigra), adult feeding, Galapagos Islands, Pacific Ocean

Galápagos tortoise or Galápagos giant tortoise (Geochelone nigra), Galapagos Islands, Pacific Ocean

Galápagos tortoise or Galápagos giant tortoise (Geochelone nigra), adult, Galapagos Islands, Pacific Ocean

Portrait of Galapagos Giant Tortoise (Geochelone elephantopus) in the Charles Darwin Station in Puerto Ayora, Insel Santa Cruz, Galapagos Inseln, Galapagos Islands, Ecuador, South America

Galapagos Giant Tortoise (Chelonoidis Nigra) Walking Through Sunlit Woods, Galapagos Islands, Ecuador

A giant Galapagos tortoise (Geochelone nigra) beds down for the night below a tree house at the Tortoise Farm on the island of Santa Cruz in the Galapagos Archipelago, Santa Cruz Island, Galapagos Islands, Ecuador