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Line of African elephants (Loxodonta africana), Samburu National Reserve, Kenya, East Africa, Africa

Lioness and cub (Panthera leo) showing affection, Masai Mara Game Reserve, Kenya, East Africa, Africa

Head on shot of lion cub (Panthera leo) looking at camera, Masai Mara Game Reserve, Kenya, East Africa, Africa

Panoramic view of trees and mountains in desert landscape, Namib Rand game reserve, Namib Naukluft Park, Namibia, Africa

Panoramic view over orange sand dunes towards mountains, Namib Rand private game reserve, Namibia, Africa

Mounted nomad holds uurga and herds horses in summer, Khogno Khan Uul Nature Reserve, Gurvanbulag, Bulgan, Northern Mongolia, Central Asia, Asia

Remains of gunpowder mills at Kennall Vale Nature Reserve in Ponsanooth near Falmouth, Cornwall, England, United Kingdom, Europe

The 659m limestone Monte Cefano Nature Reserve and Custonaci marble quarries, on the coastline north east of Trapani, Custonaci, Sicily, Italy, Mediterranean, Europe

Spring flowers and 659m limestone Monte Cefano, a Nature Reserve and hiking and climbing spot northeast of Trapani, Custonaci, Sicily, Italy, Mediterranean, Europe

Spring flowers and 659m limestone Monte Cefano, a Nature Reserve and hiking and climbing spot northeast of Trapani, Custonaci, Sicily, Italy, Mediterranean, Europe

Spring flowers in Zingaro Nature Reserve near Scopello on this north west coast, known for its beauty, Scopello, Sicily, Italy, Mediterranean, Europe

The 659m limestone Monte Cefano, Nature Reserve and hiking and climbing spot, juts into the sea northeast of Trapani, Custonaci, Sicily, Italy, Mediterranean, Europe

African elephant (Loxodonta africana) going to the Uaso Nyro River, Samburu National Reserve, Kenya, East Africa, Africa

Sheep, Hermaness Nature Reserve, Hermaness, Unst, Shetland Islands, Scotland, United Kingdom, Europe

Loch Druidibeg Nature Reserve at sunset, with Hecla, Ben Corodale and Beinn Mhor behind, South Uist, Outer Hebrides, Scotland, United Kingdom, Europe

Male Eastern yellow-billed hornbill (Tockus flavirostris), Samburu National Reserve, Kenya, East Africa, Africa

Lion (Panthera leo) carrying a blue wildebeest (brindled gnu) (Connochaetes taurinus) carcass, Masai Mara National Reserve, Kenya, East Africa, Africa

Grant•À?s zebra (Plains zebra) (common zebra) (Equus burchelli boehmi) and topi (tsessebe) (Damaliscus lunatus) crossing the Mara River, Masai Mara National Reserve, Kenya, East Africa, Africa

Red-billed oxpecker (Buphagus erythrorhynchus) and Cape buffalo (African buffalo) (Syncerus caffer), Hluhluwe Game Reserve, South Africa, Africa

Cheetah (Acinonyx jubatus) mother and two cubs, Masai Mara National Reserve, Kenya, East Africa, Africa

Cheetah (Acinonyx jubatus) on Land Rover safari vehicle, Masai Mara National Reserve, Kenya, East Africa, Africa

The altiplano at an altitude of over 4000m looking over the salt lake Laguna de Tuyajto, Los Flamencos National Reserve, Atacama Desert, Antofagasta Region, Norte Grande, Chile, South America

The altiplano at an altitude of over 4000m and the peak of Cerro Miniques at 5910m, Los Flamencos National Reserve, Atacama Desert, Antofagasta Region, Norte Grande, Chile, South America

Nile crocodile (Crocodylus niliticus) on shore of Mara River with open jaws, Masai Mara National Reserve, Kenya, East Africa, Africa

Burchell's zebras (Equus burchelli) and topi (Damaliscus korrigum) , Masai Mara National Reserve, Kenya, East Africa, Africa

Line of African elephants (Loxodonta africana), Samburu National Reserve, Kenya, East Africa, Africa

Male gerenuk (Litocranius walleri) feeding while standing on its hind legs, Samburu National Reserve, Kenya, East Africa, Africa

The magnificent and mysterious Wistmans Wood Nature Reserve in Dartmoor National Park, Devon, England, United Kingdom, Europe

Sand dunes at Murlough Nature Reserve, with views to Dundrum Bay, County Down, Ulster, Northern Ireland, United Kingdom, Europe

Ferns growing amongst the sand dunes at Murlough Nature Reserve, with views to Dundrum Bay and the Mountains of Mourne beyond, County Down, Ulster, Northern Ireland, United Kingdom, Europe

Sand dunes at Murlough Nature Reserve, with views to Dundrum Bay and the Mountains of Mourne beyond, County Down, Ulster, Northern Ireland, United Kingdom, Europe

Boats moored at Tofino, Pacific Rim National Park Reserve, Vancouver Island, British Columbia, Canada, North America

The River Fowey flowing though Draynes Wood at Golitha Falls National Nature Reserve in autumn, Cornwall, England, United Kingdom, Europe

Female Sumatran rhino (Borneo rhino) (Dicerorhinus sumatrensis) in wallow, Tabin Reserve, Sabah, Borneo, Malaysia, Southeast Asia, Asia

Female Sumatran rhino (Borneo rhino) (Dicerorhinus sumatrensis) in wallow, Tabin Reserve, Sabah, Borneo, Malaysia, Southeast Asia, Asia

Macaranga sp. an early coloniser of logged forest because its seeds are capable of germinating in light sunny clearings, Tabin Wildlife Reserve, Sarawak, Borneo, Malaysia, Southeast Asia, Asia

Chacma baboon (Papio ursinus) with its cheeks full of food, Hluhluwe Game Reserve, South Africa, Africa

Giant fruit of coco de mer palm (Lodoicea maldivica) in the Vallee de Mai Nature Reserve, UNESCO World Heritage Site, Baie Sainte Anne district, Island of Praslin, Seychelles, Africa

Birdwatcher beneath giant palm leaves in the Vallee de Mai Nature Reserve, UNESCO World Heritage Site, Baie Sainte Anne district, Island of Praslin, Seychelles, Africa

Male Spotted-backed weaver (Village weaver) (Ploceus cucullatus) building a nest, Hluhluwe Game Reserve, South Africa, Africa

Tourist walking down the steps to Calamosche Beach, near Noto, Vendicari Nature Reserve, South East Sicily, Italy, Mediterranean, Europe

Calamosche Beach, a popular, secluded beach near Noto, Vendicari Nature Reserve, South East Sicily, Italy, Mediterranean, Europe

The Namaqua chameleon (Chamaeleo namaquensis) on man's hand, NamibRand Nature Reserve, Namib Desert, Namibia, Africa

The sun starts to light up the distant mountains in the NamibRand Nature Reserve, Namib Desert, Namibia, Africa

A large herd of Gemsbok (Oryx gazella) on the plains of the NamibRand Nature Reserve, Namib Desert, Namibia, Africa

Storm clouds and rainbow in the early evening in NamibRand Nature Reserve, Namib Desert, Namibia, Africa

Gemsbok (Oryx gazella) sheltering from the midday sun in the NamibRand Nature Reserve, Namib Desert, Namibia, Africa

Alligator (Yacare caiman), Pantanal Conservation Area, UNESCO World Heritage Site, Brazil, South America

Wild garlic (ramsons) (Allium ursinum) growing in Black Woods nature reserve in Cheddar Gorge, Somerset, England, United Kingdom, Europe

Sunrise over a meadow of snake's head fritillary (Fritillaria meleagris) wildflowers growing at North Meadow National Nature Reserve, Cricklade, Wiltshire, England, United Kingdom, Europe

Seventy Islands (Ngerukewid Islands Wildlife Preserve), forest-covered limestone rock, protected as a Nature Reserve, so can only be seen from the air, Palau, Micronesia, Western Pacific Ocean, Pacific

Seventy Islands (Ngerukewid Islands Wildlife Preserve), forest-covered limestone rock, protected as a Nature Reserve, so can only be seen from the air, Palau, Micronesia, Western Pacific Ocean, Pacific

Seventy Islands (Ngerukewid Islands Wildlife Preserve), forest-covered limestone rock, protected as a Nature Reserve, so can only be seen from the air, Palau, Micronesia, Western Pacific Ocean, Pacific

Four Masai hunters running through bush holding spears, Masai Mara National Reserve, Kenya, East Africa, Africa