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Cape buffalo (African buffalo) (Syncerus caffer) and two yellow-billed oxpeckers (Buphagus africanus), Ngorongoro Crater, Tanzania, East Africa, Africa

Cape Buffalo (African buffalo) (Syncerus caffer), white-naped raven (white-necked raven) (Corvus albicollis) and a yellow-billed oxpecker (Buphagus africanus), Ngorongoro Crater, Tanzania, East Africa, Africa

Redbilled oxpecker (Buphagus erythrorhynchus), on giraffe, Kruger National Park, South Africa, Africa

Sugar cane harvest, Ramadi village, west bank of the Nile south of Edfu, Egypt, North Africa, Africa

Adult black-browed albatross (Thalassarche melanophrys), pair in courtship display at nesting site on New Island, Falklands, South America

Racehorses and jockeys going over the sticks at Cheltenham Racecourse for the National Hunt Festival of Racing, UK

Alaskan Huskies harnessed for dog-sledding at Villmarkssenter wilderness centre, Kvaloya Island, Tromso in Arctic Circle Northern Norway

Racehorses and jockeys going over the sticks at Cheltenham Racecourse for the National Hunt Festival of Racing, UK

Alaskan Huskies dog-sledding at Villmarkssenter wilderness centre on Kvaloya Island, Tromso in Arctic Circle Northern Norway

Alaskan Huskies dog-sledding at Villmarkssenter wilderness centre on Kvaloya Island, Tromso in Arctic Circle, Northern Norway

Hungarian Csikos cowboy giving display of horsemanship skills on The Great Plain of Hungary at Bugac, Hungary

Alaskan Huskies dog-sledding at Villmarkssenter wilderness centre on Kvaloya Island, Tromso in Arctic Circle, Northern Norway

Pair of Weavers birds on tree branch at Chattra Sagar nature reserve at Nimaj, Rajasthan, Northern India

Alaskan Huskies dog-sledding at Villmarkssenter wilderness centre on Kvaloya Island, Tromso in Arctic Circle Northern Norway

Alaskan Huskies dog-sledding at Villmarkssenter wilderness centre on Kvaloya Island, Tromso in Arctic Circle, Northern Norway

Alaskan Huskies dog-sledding at Villmarkssenter wilderness centre on Kvaloya Island, Tromso in Arctic Circle Northern Norway

Racehorses and jockeys going over the sticks at Cheltenham Racecourse for the National Hunt Festival of Racing, UK

Hungarian Csikos cowboy showing horsemanship skills with horse on The Great Plain of Hungary at Bugac, Hungary

Alaskan Huskies dog-sledding at Villmarkssenter wilderness centre on Kvaloya Island, Tromso in Arctic Circle, Northern Norway

Alaskan Huskies harnessed for dog-sledding at Villmarkssenter wilderness centre Kvaloya Island, Tromso, Arctic Circle Northern Norway

Alaskan Huskies eagerly wait at Villmarkssenter wilderness adventure centre on Kvaloya Island, Tromso in Arctic Circle Northern Norway

Alaskan Huskies dog-sledding at Villmarkssenter wilderness centre on Kvaloya Island, Tromso in Arctic Circle Northern Norway

Alaskan Huskies dog-sledding at Villmarkssenter wilderness centre on Kvaloya Island, Tromso in Arctic Circle, Northern Norway

Hungarian Csikos cowboy giving display of horsemanship skills on The Great Plain of Hungary at Bugac, Hungary

Racehorses and jockeys going over the sticks at Cheltenham Racecourse for the National Hunt Festival of Racing, UK

Hugarian Csikos - Cowboy - with his horse resting his hoof on his leg at Bugac on the Great Plain of Hungary

Racehorses and jockeys going over the sticks at Cheltenham Racecourse for the National Hunt Festival of Racing, UK

Racehorses and jockeys going over the sticks at Cheltenham Racecourse for the National Hunt Festival of Racing, UK

Racehorses and jockeys on the home straight at Cheltenham Racecourse for the National Hunt Festival of Racing, UK

Interior, Tower Bridge House, architects Richard Rogers Partnership, Tower Hamlets, London, England, United Kingdom, Europe

Main chamber, National Assembly for Wales, architect Richard Rogers Partnership, Cardiff, Wales, United Kingdom, Europe

National Assembly for Wales, architects Richard Rogers Partnership, Cardiff, Wales, United Kingdom, Europe

LutherÌs Partner Goby (Cryptocentrus lutheri) Lives in partnership with the snapping shrimp Alpheus djiboutensis. One of many Goby/Shrimp partnerships, The goby always choosing to pair with the same species of shrimp. The goby keeps a watchful eye for the almost blind shrimp, in return for sanctuary in the hosts burrow. The shrimp digs constantly and maintains contact with the goby at all times via antennae. Red Sea.

Harbour porpoise (Phocoena phocoena). A mother and calf pair, the mother showing that photo id of porpoises may well be possible. Hebrides, Scotland

Fraser's dolphin (Lagenodelphis hosei). A bow riding Frase's dolphin showing clearly the distinctive pink colouring.Caribbean.

An RAF Sea King Helicopter and search dog and handler, Lake District, Cumbria, England, United Kingdom, Europe

Eurasian river otter (Lutra lutra) mother eating fish with cub closeby. The cub tried unsuccessfully to obtain some of its mother's catch. Hebrides, Scotland

Blue-footed booby (Sula nebouxii) in the Galapagos Island Group, Ecuador. The Galapagos are a nest and breeding area for blue-footed boobies.

American oystercatcher (Haematopus ostralegus) along the shoreline on Bartolome Island in the Galapagos Island Group, Ecuador. Pacific Ocean.

Breeding adult male and female harlequin ducks (Histrionicus histrionicus) in Red Bluff Bay on Baranof Island, Southeast Alaska

Fighting adult Royal Penguins (Eudyptes Schlegeli). Sandy Bay, Macquarie Island, Subantarctic Australia.

Eurasian river otter (Lutra lutra) mother and cub. Cubs often have darker fur than their mothers and for the first few months are much more 'buoyant' in the water. Hebrides, Scotland

Sanderling (Calidris alba) running from incoming waves early in the morning silhouetted against the orange early light. Soroby. Argyll,. Scotland, UK

Great frigate bird (Fregata minor) nesting and breeding site on North Seymour Island in the Galapagos Island Group, Ecuador. Pacific Ocean.

Two bottlenose dolphins (Tursiops truncatus) and their shadows in the shallows of the Caicos Bank. Turks and Caicos Islands. (A4 Only)

Short finned pilot whale (Globicephala macrorynchus). A young pilot whale hovers at the surface near to its mother. Gulf of California.