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Adult Atlantic puffins (Fratercula arctica), breeding on the island of Bolscheoya in the Svalbard Archipelago, Norway

Adult Atlantic puffin (Fratercula arctica), returning to the nest site with fish (sand eels) for its chick on Grimsey Island, Iceland, Polar Regions

Adult Atlantic puffin (Fratercula arctica), returning to the nest site with fish (sand eels) for its chick on Grimsey Island, Iceland, Polar Regions

Adult Atlantic puffin (Fratercula arctica), returning to the nest site with fish (sand eels) for its chick on Grimsey Island, Iceland, Polar Regions

Adult Atlantic puffins (Fratercula arctica), gathering above the nest site on Grimsey Island, Iceland, Polar Regions

Adult Atlantic puffins (Fratercula arctica), returning to the nest site with fish (sand eels) for its chick on Grimsey Island, Iceland, Polar Regions

Adult puffin (Fratercula arctica) taking flight near the nesting island of Vaeroya in the Lofoton Island Group of Northern Norway, Norwegian Sea.

Adult puffin (Fratercula arctica) in flight near the nesting island of Vaeroya in the Lofoton Island Group of Northern Norway, Norwegian Sea.

Adult puffin (Fratercula arctica) taking flight near the nesting island of Vaeroya in the Lofoton Island Group of Northern Norway, Norwegian Sea.

Adult puffin (Fratercula arctica) taking flight on the nesting island of Vaeroya in the Lofoton Island Group of Northern Norway, Norwegian Sea. (rr)

Atlantic puffin (Fratercula arctica) on sea cliff top and flying gannets in seabird colony, Shetland Islands, Scotland, UK

Atlantic puffin (Fratercula arctica) on sea cliff top watching gannet flying by in seabird colony, Scotland, UK

Atlantic puffin (Fratercula arctica) on sea cliff top and great skua chasing gannet in seabird colony, Shetland Islands, Scotland, UK

Two Puffins (Fratercula arctica), standing in the grass, bird rock Latrabjard, Westfjords, Iceland, Europe

Puffin (Fratercula arctica), standing in the grass, bird rock Latrabjard, Westfjords, Iceland, Europe

Puffin (Fratercula arctica), standing on grass, calling, bird rock Latrabjard, Westfjords, Iceland, Europe

Atlantic puffin (Fratercula arctica) on sea cliff top and flying gannets in seabird colony, Shetland Islands, Scotland, UK

Puffin (Fratercula arctica), standing in the grass, bird rock Latrabjard, Westfjords, Iceland, Europe

Puffin (Fratercula arctica), Puffin on Bird Rock, Staple Island, Farne Islands Nature Reserve, Farne Islands, Northumberland, England, United Kingdom, Europe

Puffin (Fratercula arctica), Puffin, nesting material in beak, Staple Island, Farne Islands Nature Reserve, Farne Islands, Northumberland, England, United Kingdom, Europe

Puffin (Fratercula arctica), standing in the grass, bird rock Latrabjard, Westfjords, Iceland, Europe

Two Puffins (Fratercula arctica), standing in the grass, bird rock Latrabjard, Westfjords, Iceland, Europe