Greater kudu (Tragelaphus strepsiceros) buck marking its territory, Addo Elephant National Park, South Africa, Africa

Lion (Panthera leo) scratching and marking territory, Serengeti National Park, Tanzania, East Africa, Africa

Cheetas scent marking their territory, Moremi, Botswana, Southern Africa

Eurasian river otter (Lutra lutra) sprainting on high point. The spraint is a method of marking territory and high points are a favoured location, even those washed by tides.

Eurasian river otter (Lutra lutra) sprainting on high point. The spraint is a method of marking territory and high points are a favoured location, even those washed by tides.

Family of Eurasian river otters (Lutra lutra) sprainting on high point. The spraint is a method of marking territory and high points are a favoured location, even those washed by tides.

Portrait of sled dog, floppy ears, Yukon Territory, Canada

Sunlit portrait of a sled dog, Yukon Territory, Canada

Brown bear (Ursus arctos) male rubbing his back on a pine tree to leave his scent, Finland

Cape Sugarbird (Promerops cafer), calling in territorial display, Western Cape, South Africa, July

Red fox (Vulpes vulpes) males in territorial confrontation. Montreal Botanical Garden. Quebec. Canada

Male Chamois marking his territory, Jura Switzerland

Cape Sugarbird (Promerops cafer), waving tail in territorial display, Western Cape, South Africa, July

Male Chamois marking his territory, Jura Switzerland

Male red deer during rutting, France

Male red deer during rutting, France

Male red deer during rutting, France