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Galapagos white-cheeked pintail (Anas bahamensis galapagensis) feeding near giant tortoise in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) with photographer at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Young Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the tortuguero breeding station, Puerto Villamil on Isabela Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Photographers with captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador