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Tombs of Ataxerxes I and Darius the Great, Naqsh-e Rostam Necropolis, near Persepolis, Iran, Middle East

Tombs of Darius II, Ataxerxes I and Darius the Great, Naqsh-e Rostam Necropolis, near Persepolis, Iran, Middle East

Carved relief of four Royal Persian Guards, facade of Private Palace of Darius the Great (Tachara), Persepolis, UNESCO World Heritage Site, Iran, Middle East

Tomb of Ataxerxes I and carved relief below, Naqsh-e Rostam Necropolis, near Persepolis, Iran, Middle East

Carved relief of lion overcoming a bull, symbolising the advent of Spring and the Iranian New Year (Nowruz), Apadana Palace, Persepolis, UNESCO World Heritage Site, Iran, Middle East

Interior of dome of Vank (Armenian) Cathedral with Archbishop's throne in foreground, Isfahan, Iran, Middle East

View across Naqsh-e (Imam) Square, UNESCO World Heritage Site, from Ali Qapu Palace opposite Sheikh Lotfollah Mosque, Isfahan, Iran, Middle East

View across Naqsh-e (Imam) Square from Ali Qapu Palace opposite Sheikh Lotfollah Mosque, UNESCO World Heritage Site, Isfahan, Iran, Middle East

Mural of Battle of Chaldoran 1518 AD, Chehel Sotun (Chehel Sotoun) (40 Columns) Palace, Isfahan, Iran, Middle East

Ceiling of The Music Hall of Ali Qapu Palace with early acoustic design, Ali Qapu Palace, Isfahan, Iran, Middle East

Mural of reception for Ruler of Turkistan, 1621 AD, Chehel Sotun (Chehel Sotoun) (40 Columns) Palace, Isfahan, Iran, Middle East

View across Naqsh-e (Imam) Square, UNESCO World Heritage Site, from Ali Qapu Palace opposite Sheikh Lotfollah Mosque, Isfahan, Iran, Middle East

Megapode hunters hunting for eggs of the Megapode bird (Megapodiidae), East New Britain, Papua New Guinea, Pacific

The Beehives (Dawapia Rocks) in Simpson Harbour, Rabaul, East New Britain, Papua New Guinea, Pacific

The Beehives (Dawapia Rocks) in Simpson Harbour, Rabaul, East New Britain, Papua New Guinea, Pacific

The Beehives (Dawapia Rocks) in Simpson Harbour, Rabaul, East New Britain, Papua New Guinea, Pacific

Tiling on facade, Aramgah-e Shah-e Cheragh (Mausoleum of the King of Light), Shiraz, Iran, Middle East

Silhouetted workman setting up marquee for special event in the courtyard of Imam Mosque, Isfahan, Iran, Middle East

Astane Square in front of the Imam Hassan Mosque, unusually challenging message (Can't Get No Satisfaction) on girl's back, Iran, Middle East

African open-billed stork (African openbill) (Anastomus lamelligerus) with a snail, Selous Game Reserve, Tanzania, East Africa, Africa

Common agama (red-headed rock agama) (rainbow agama) (Agama agama), male, Ruaha National Park, Tanzania, East Africa, Africa

White-headed buffalo weaver (Dinemellia dinemelli), Serengeti National Park, Tanzania, East Africa, Africa

Two male lion (Panthera leo) attacking a Cape Buffalo (African Buffalo) (Syncerus caffer), Ruaha National Park, Tanzania, East Africa, Africa

White-crowned lapwing (white-headed lapwing) (white-headed plover) (white-crowned plover) (Vanellus albiceps), Selous Game Reserve, Tanzania, East Africa, Africa

Von Der Decken's hornbill (Tockus deckeni), male, Selous Game Reserve, Tanzania, East Africa, Africa

Madagascar squacco heron (Madagascar pond-heron) (Malagasy pond-heron) (Ardeola idae), Selous Game Reserve, Tanzania, East Africa, Africa

Yellow baboon (Papio cynocephalus) juvenile eating a duom palm fruit, Selous Game Reserve, Tanzania, East Africa, Africa

Angola colobus (Angolan black-and-white colobus) (Angolan colobus) (Colobus angolensis), Selous Game Reserve, Tanzania, East Africa, Africa

Red-necked spurfowl (red-necked francolin) (Francolinus afer) (Pternistes afer), Mikumi National Park, Tanzania, East Africa, Africa

Red-billed oxpecker (Buphagus erythrorhynchus), Ngorongoro Conservation Area, Tanzania, East Africa, Africa

Saddle-billed stork (Ephippiorhynchus senegalensis), Ngorongoro Crater, Tanzania, East Africa, Africa

Mwanza flat-headed rock agama (Agama mwanzae), Serengeti National Park, Tanzania, East Africa, Africa

Portrait of a female cheetah (Acinonyx jubatus) in tall grass, Samburu National Reserve, Kenya, East Africa, Africa

A Grant's zebra (Equus quagga boehmi) looking at the camera, Masai Mara National Reserve, Kenya, East Africa, Africa

Tree Squirrel (Smith's bush squirrel) (yellow-footed squirrel) (Paraxerus cepapi), Ruaha National Park, Tanzania, East Africa, Africa