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Swainson's Francolin (Spurfowl) (Pternistes swainsonii), Kruger National Park, South Africa, Africa

Rhinoceros and calf at Ziwa Rhino Sanctuary, Uganda, East Africa, Africa

Cruise Ship off the coast of Mykonos Island, Cyclades, Greek Islands, Greece, Europe

Mountain hare (Lepus timidus) in winter snow, Scottish Highlands, Scotland, United Kingdom, Europe

Profile of American Bison (Bison Bison) in the snow in the Badlands, Badlands National Park, South Dakota, United States

Close-up of head of Chinstrap penguin (Pygoscelis antarcticus), Half Moon Island, Antarctica, Polar Regions

Hoodoo spires rising above the valley below the south side of Blue Mesa in Petrified Forest National Park, Arizona, United States of America, North America

Alpaca in Ollantaytambo, Peru, South America

Frozen bison (Bison bison), sticking out tongue, Yellowstone National Park, UNESCO World Heritage Site, Wyoming, United States of America, North America

Crimson-breasted boubou (crimson-breasted shrike) (Laniarius atrococcieneus), Kgalagadi Transfrontier Park, encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Two springbok (Antidorcas marsupialis) bucks fighting, Kgalagadi Transfrontier Park, encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Cape mountain zebra (Equus zebra zebra), Karoo National Park, South Africa, Africa

Tour boats in Kleftiko Bay, white cliffs of Kleftiko, Milos, Cyclades Islands, Greek Islands, Aegean Sea, Greece, Europe

Loggerhead turtle (Caretta caretta) hatchling, moving from nest to sea at night, Banga Nek, Kwazulu Natal, South Africa, Africa

Moose (Alces alces) cow, Glacier National Park, Montana, United States of America, North America

Fur seal (Arctocephalus forsteri), Moeraki, South Island, New Zealand, Pacific

Black bear (Ursus americanus), Waterton Lakes National Park, Alberta, Canada, North America

Oystercatcher, County Clare, Munster, Republic of Ireland, Europe

Chough, County Clare, Munster, Republic of Ireland, Europe

Red panther chameleon (Furcifer pardalis), endemic to Madagascar, Africa

Sanderling (Calidris alba), County Clare, Munster, Republic of Ireland, Europe

Double-crested cormorant (Phalacrocorax auritus), Sonny Bono Salton Sea National Wildlife Refuge, California, United States of America, North America

Female rufous hummingbird (Selasphorus rufus) perched, Routt National Forest, Colorado, United States of America, North America

Red grouse (Lagopus lagopus), male, in heather, County Durham, England, United Kingdom, Europe

Basilica of St. Maria del Popolo, Lion of Fountain of Obelisk, Piazza del Popolo, UNESCO World Heritage Site, Rome, Latium (Lazio), Italy, Europe

Lion of Fountain of Obelisk, Piazza del Popolo, UNESCO World Heritage Site, Rome, Latium (Lazio), Italy, Europe

Beaver (Castor canadensis) swimming with food, Denali National Park and Preserve, Alaska, United States of America, North America

Red squirrel (spruce squirrel) (Tamiasciurus hudsonicus) in a spruce tree, Denali National Park and Preserve, Alaska, United States of America, North America

Leopard at night (Panthera pardus), South Luangwa National Park, Zambia, Africa

Great frigate bird (Frigata minor), Isla Genovesa, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Pronghorn (Antilocapra americana) in the snow, Grand Teton National Park, Wyoming, United States of America, North America

Tawny eagle (Aquila rapax), Lualenyi Game Reserve, Kenya, East Africa, Africa

Elephants (Loxodonta africana), Masai Mara National Reserve, Kenya, East Africa, Africa

A single wild brown horse stands in a snowy landscape, United Kingdom, Europe

Woman in colourful sari with a painted ceremonial elephant in Jaipur, Rajasthan, India, Asia

Moai heads of Easter Island, Rapa Nui National Park, UNESCO World Heritage Site, Easter Island, Chile, South America

Clymene dolphin (Stenella clymene) porpoising with water trailing its flanks, offshore Senegal, West Africa, Africa

Roman road, Temple of Castor and Pollux, Palatine Hill behind, Roman Forum, UNESCO World Heritage Site, Rome, Lazio, Italy, Europe

A young dusky leaf monkey (spectacled langur) (dusky langur) in Langkawi rainforest, Malaysia, Southeast Asia, Asia

View of the Campanile and Doge's Palace in St. Mark's Square, Venice, UNESCO World Heritage Site, Veneto, Italy, Europe

Nyhavn with old colourful buildings and boats anchored in summer, Copenhagen, Denmark, Europe

Bull elk (Cervus canadensis) eating yellow wildflowers, Jasper National Park, Alberta, Canada, North America

Traditionally dressed huaso riding horse on field, Colina, Chacabuco Province, Santiago Metropolitan Region, Chile, South America

Huaso riding horse at ranch on sunny day, Colina, Chacabuco Province, Santiago Metropolitan Region, Chile, South America

An ornate colourful door at Eng Chuan Tong Tan Kongsi clan house in George Town, Penang Island, Malaysia, Southeast Asia, Asia

Green Plumed Basilisk Lizard (Basiliscus plumifrons), Boca Tapada, Alajuela Province, Costa Rica, Central America

Moai heads of Easter Island, Rapa Nui National Park, UNESCO World Heritage Site, Easter Island, Chile, South America

Brace of Atlantic Puffins, United Kingdom, Europe

Pont du Gard, Roman aqueduct, UNESCO World Heritage Site, River Gard, Languedoc-Roussillon, southern France, France, Europe

Adult male proboscis monkey (Nasalis larvatus), endemic to Borneo, Tanjung Puting National Park, Borneo, Indonesia, Southeast Asia, Asia

Southern red-billed hornbill (Tockus rufirostris), Kruger National Park, South Africa, Africa

Common waterbuck (Ellipsen waterbuck) (Kobus ellipsiprymnus ellipsiprymnus), doe and calf, Kruger National Park, South Africa, Africa

European roller (Coracias garrulus), Kruger National Park, South Africa, Africa

Weikersheim Castle, Hohenlohe Region, Taubertal Valley, Romantische Strasse (Romantic Road), Baden Wurttemberg, Germany, Europe

Rock formations in the foothills of Andes in Cafayate region, Salta, Argentina, South America

Photographer in the Sierra de San Francisco, El Vizcaino Biosphere Reserve, Baja California Sur, Mexico, North America

Adult white-faced ibis (Plegadis chihi), foraging in tidal estuary, San Jose del Cabo, Baja California Sur, Mexico, North America

A view of Liberty Bridge and the Pest side, Budapest, Hungary, Europe

A dancer performing at the Melia Rio de Oro Hotel, Playa Esmeralda, Guardalavaca, Cuba, West Indies, Central America

Long-tailed macaques (Macaca fascicularis)grooming near Angkor Thom, Siem Reap, Cambodia, Indochina, Southeast Asia, Asia

First Nations totem poles, Thunderbird Park, Vancouver Island, next to the Royal British Columbia Museum, Victoria, British Columbia, Canada, North America

Young African bush elephants (Loxodonta africana), playing in the water, Tarangire National Park, Tanzania, East Africa, Africa

African bush elephant (Loxodonta africana), mother and calves, Tarangire National Park, Tanzania, East Africa, Africa

Detail from the Column of Trajan, Trajan Forum (Foro Traiano), UNESCO World Heritage Site, Rome, Lazio, Italy, Europe

Cheetah (Acinonyx jubatus) cub, Ngorongoro Conservation Area, UNESCO World Heritage Site, Serengeti, Tanzania, East Africa, Africa

Yellow billed hornbill, Botswana, Africa

Arctic tern (Sterna paradisaea) with sand eel, Inner Farne, Farne Islands, Northumberland, England, United Kingdom, Europe

Cape buffalo (Syncerus caffer) herd, Zimanga private game reserve, KwaZulu-Natal, South Africa, Africa

Blackbacked jackal (Canis mesomelas) with whitebacked vultures (Gyps africanus), Zimanga private game reserve, KwaZulu-Natal, South Africa, Africa

Sable (Hippotragus niger), Chobe National Park, Botswana, Africa

Lizard in the forest next to the Iguazu Falls, Foz do Iguacu, State of Parana, Brazil, South America

Black-browed albatross (Thalassarche melanophris) in breeding colony on Saunders Island, Falkland Islands, South America

Indian thick-knee (Burhinus indicus), Bandhavgarh National Park, Madhya Pradesh, India, Asia

A grey-headed kingfisher (Halcyon leucocephala) perched on a tree branch, Samburu, National Reserve, Kenya, East Africa, Africa

Three king penguins (Aptenodytes patagonica) walking on Volunteer Point beach, Falkland Islands, South America

Ponte Sant' Angelo and Castel Sant'Angelo, Vatican area, Historic Centre, Rome, UNESCO World Heritage Site, Lazio, Italy, Europe

Castel Sant'Angelo (Hadrian's Mausoleum), Vatican area, Historic Centre, Rome, UNESCO World Heritage Site, Lazio, Italy, Europe

St. Pauls Cathedral, London, England, United Kingdom, Europe

Woman looks at old church perched on mountains, San Romerio Alp, Brusio, Canton of Graubunden, Poschiavo Valley, Switzerland, Europe

An adult blackspotted puffer (Arothron nigropunctatus), off Bangka Island, near Manado, Sulawesi, Indonesia, Southeast Asia, Asia

Rows of empty beach loungers in Juan les Pins, Cote d'Azur, Provence, France, Mediterranean, Europe

The colorful sea village of Riomaggiore, Cinque Terre, UNESCO World Heritage Site, Liguria, Italy, Europe

Hiker illuminated by a beam of sunlight after snowfall in the Denali National Park, Alaska, United States of America, North America

Wild dingo on Seventy Five Mile Beach, Fraser Island, Queensland, Australia, Pacific

Wood boat on the shore of River Mera at sunrise, Sorico, Como province, Lower Valtellina, Lombardy, Italy, Europe

Swans in River Mera at sunrise, Sorico, Como province, Lower Valtellina, Lombardy, Italy, Europe

White rhinoceros (Ceratotherium simum), Kalahari, Botswana, Africa

Lion (Panthera leo), Savuti, Chobe National Park, Botswana, Africa

Man walking under a lone palm on the beach at Soufriere with Petit Piton in the distance, St. Lucia, Windward Islands, West Indies Caribbean, Central America

Hippo (Hippopotamus amphibius), Chobe River, Botswana, Africa

Hippo (Hippopotamus amphibius) yawning, Chobe National Park, Botswana, Africa

St. Peter's Basilica (Basilica di San Pietro), St. Peter's Square (Piazza de San Pietro), UNESCO World Heritage Site, Vatican City, Rome, Lazio, Italy, Europe