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Wild Galapagos giant tortoise (Geochelone elephantopus), with tourists on Santa Cruz Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Preserved Galapagos giant tortoise (Chelonoidis spp), known as Lonesome George, Santa Cruz Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Geochelone elephantopus vandenburghi, Giant Tortoise Breeding Centre, Puerto Villamil, Isla Isabela, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Wild Galapagos giant tortoise (Geochelone elephantopus) at Urbina Bay, Isabela Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) with photographer at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Galapagos white-cheeked pintail (Anas bahamensis galapagensis) feeding near giant tortoise in the Galapagos Island Archipelago, Ecuador

Young Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the tortuguero breeding station, Puerto Villamil on Isabela Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Group, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Group, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Photographers with captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus) at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Two adult captive Galapagos giant tortoise (Geochelone elephantopus), Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) feeding on fallen passion fruit on the upslope grasslands of Santa Cruz Island in the Galapagos Island Archipelago, Ecuador

Captive Galapagos giant tortoise (Geochelone elephantopus) being fed at the Charles Darwin Research Station on Santa Cruz Island in the Galapagos Island Group, Ecuador

Wild Galapagos giant tortoise (Geochelone elephantopus) on the upslope grasslands of Cruz Island in the Galapagos Island Group, Ecuador

Galápagos tortoise or Galápagos giant tortoise (Geochelone nigra), adult feeding, Galapagos Islands, Pacific Ocean

Galápagos tortoise or Galápagos giant tortoise (Geochelone nigra), Galapagos Islands, Pacific Ocean

Galápagos tortoise or Galápagos giant tortoise (Geochelone nigra), adult, Galapagos Islands, Pacific Ocean

Aldabra Giant Tortoise (Aldabrachelys gigantea, syn. Geochelone gigantea, Dipsochelys elephantina and Dipsochelys dussumieri), portrait, Curieuse island, Praslin, Seychelles, Africa, Indian Ocean

Aldabra giant tortoise (Geochelone gigantea), Aldabra Giant Tortoise, Granite Island, La Digue, Seychelles, La Digue, Seychelles, Africa

Portrait of Galapagos Giant Tortoise (Geochelone elephantopus) in the Charles Darwin Station in Puerto Ayora, Insel Santa Cruz, Galapagos Inseln, Galapagos Islands, Ecuador, South America

Tourists converge around a captive giant tortoise housed at the Darwin Research Station on Santa Cruz Island, Galapagos.

Galapagos giant tortoise (Chelonoidis nigra) lumbers slowly across a long, straight dirt road that stretches off to the horizon. Beyond the grass verge, there is dense forest on either side; Galapagos Islands, Ecuador

Galapagos giant tortoise (Chelonoidis nigra) lumbers slowly across a long, straight dirt road that stretches off to the horizon. Beyond the grass verge, there is dense forest on either side; Galapagos Islands, Ecuador

Galapagos Giant Tortoise (Geochelone Hoodensis) In Artificial Concrete Pool, Galapagos Islands, Ecuador

Galapagos Giant Tortoise (Chelonoidis Nigra) Walking Through Sunlit Woods, Galapagos Islands, Ecuador

Galapagos Giant Tortoise (Geochelone Hoodensis) In Artificial Concrete Pool, Galapagos Islands, Ecuador