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Moulting adult king penguins, Aptenodytes patagonicus, in the snow at Grytviken, South Georgia Island, Atlantic Ocean

Colourful reef fishes on the reef at Tumakohua Pass, Fakarava Atoll, Tuamotus, French Polynesia, South Pacific, Pacific

Resident killer whale, Orcinus orca, tail throw in Chatham Strait, Southeast Alaska, United States of America

An adult sungrebe, Heliornis fulica, swimming in Cano Chiquerra, Tortuguero National Park, Costa Rica, Central America

A juvenile male common basilisk, Basiliscus basiliscus, in Tortuguero National Park, Costa Rica, Central America

An adult crab-eating raccoon, Procyon cancrivorus, Manuel Antonio National Park, Costa Rica, Central America

Whale shark (Rhincodon typus) vertical suction feeding on a shoal of red fish, Honda Bay, Palawan, The Philippines, Southeast Asia, Asia

Whale shark (Rhincodon typus) surrounded by a shoal of fish evading predation, Honda Bay, Palawan, The Philippines, Southeast Asia, Asia

White-backed vulture (Gyps africanus) dominance display, Zimanga private game reserve, KwaZulu-Natal, South Africa, Africa

Two short-beaked common dolphins (Delphinus delphis) surfacing, Northeast Atlantic, offshore Morocco, North Africa, Africa

White rhino (Ceratotherium simum) bulls in confrontation, Zimanga private game reserve, KwaZulu-Natal, South Africa, Africa

Ground squirrels (Geosciurus inauris), Kgalagadi Transfrontier Park, Northern Cape, South Africa, Africa

Nazca Booby (Sula granti) with open mouth, Genovesa Island, Galapagos National Park, Ecuador, South America

Breaching humpback whale (Megaptera novaeangliae), Hawaii, United States of America, Pacific, North America

A black and yellow ornate ghost pipefish (Solenostomus paradoxus) pictured against coral, Russell Islands, Solomon Islands, South Pacific, Pacific

Red Fox (Vulpes vulpes) (Vulpes fulva) vixen and two kits, Yellowstone National Park, Wyoming, United States of America, North America

Young Common Waterbuck (Ellipsen Waterbuck) (Kobus ellipsiprymnus), Kruger National Park, South Africa, Africa

Pronghorn (Antilocapra americana) buck, Yellowstone National Park, Wyoming, United States of America, North America

Terns and seagulls on Capitola Beach, Capitola City, Santa Cruz County, California, United States of America, North America

Adult yellow-crowned night heron (Nyctanassa violacea), Puerto Egas, Santiago Island, Galapagos Islands, Ecuador, South America

Secretarybird (Sagittarius serpentarius), Kgalagadi Transfrontier Park, encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Two gemsbok (South African oryx) (Oryx gazella) fighting, Kgalagadi Transfrontier Park, encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Lion (Panthera leo) pair, Kgalagadi Transfrontier Park, encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Greater kudu (Tragelaphus strepsiceros) buck, Kgalagadi Transfrontier Park, encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Adult bull Antarctic fur seal, Arctocephalus gazella, head detail at Ocean Harbour, South Georgia Island, Atlantic Ocean

Rosy-breasted longclaw (pink-throated longclaw) (Macronyx ameliae) singing, Ngorongoro Crater, Tanzania, East Africa, Africa

Canadian Lynx (Lynx canadensis) in snow in captivity, near Bozeman, Montana, United States of America, North America

An adult striated caracara, Phalcoboenus australis, harassing a gentoo penguin, New Island, Falkland Islands, South Atlantic Ocean

Kittiwake (Rissa tridactyla) perched on the edge of a ledge against a backdrop of cliffs at Bempton, Yorkshire, England, United Kingdom, Europe

Male Impala (Aepyceros melampus) with a Red-Billed Oxpecker (Buphagus erythrorhynchus), Kruger National Park, South Africa, Africa

Grey crowned crane (Southern crowned crane) (Balearica regulorum), Serengeti National Park, Tanzania, East Africa, Africa

Red-billed oxpecker (Buphagus erythrorhynchus) on a zebra, Imfolozi Game Reserve, South Africa, Africa

Gray wolf (Canis lupus) 755M of the Lamar Canyon Pack running through the snow in the winter, Yellowstone National Park, Wyoming, United States of America, North America

Resident killer whale, Orcinus orca, Cattle Pass, San Juan Island, Washington, United States of America, North America

Shipwreck off the Atlantic coast, shot with long exposure to record motion in sea and sky, near Walvis Bay, Namibia, Africa

Flamingos at Red Lagoon (Laguna Colorada), a salt lake in the Altiplano of Bolivia in Eduardo Avaroa Andean Fauna National Reserve, Bolivia, South America

Close-up of mantle of magnificent anemone (Heteractis magnifica), Ras Mohammed National Park, off Sharm el-Sheikh, Sinai, Red Sea, Egypt, North Africa, Africa

Silhouette of two scuba divers above coral reef scene, Ras Mohammed National Park, Sharm el-Sheikh, Red Sea, Egypt, North Africa, Africa

Giant trevally (Caranx ignobilis) shoal schooling, Ras Mohammed National Park, Red Sea, Egypt, North Africa, Africa

Snowy Egret (Egretta thula) by the Nosara River mouth near the Biological Reserve, Nosara, Guanacaste Province, Costa Rica, Central America

Close up of a male Southern elephant seal (Mirounga leonina), St. Andrews Bay, South Georgia Island, Polar Regions

Adult polar bear (Ursus maritimus) on the ice in Bear Sound, Spitsbergen Island, Svalbard, Norway, Scandinavia, Europe

Harbour seal (common seal) (Phoca vitulina), Foula Island, Shetlands, Scotland, United Kingdom, Europe

Three bald eagle (Haliaeetus leucocephalus) in an evergreen tree, Yellowstone National Park, Wyoming, United States of America, North America

Majestic stag with large antlers standing in a sunlit field, showcasing wildlife in a natural habitat, United Kingdom, Europe

Marine Iguana (Amblyrhynchus cristatus), Turtle Bay, Isla Santa Cruz, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Pronghorn (Antilocapra americana) buck, Custer State Park, South Dakota, United States of America, North America

Giant sponge showing how it filters water with the use of dye, Dominica, West Indies, Caribbean, Central America

A rare longnose walking batfish (Ogcocephalus corniger) that usually lives at depths to 300m, Dominica, West Indies, Caribbean, Central America

Polar bear (Ursus maritimus) and cub, Wapusk National Park, Churchill, Hudson Bay, Manitoba, Canada, North America

Four mountain gorillas (Gorilla gorilla beringei) of the Amahoro group, Volcanoes National Park, Rwanda, Africa

Blue wildebeest (brindled gnu) (Connochaetes taurinus) herd, Serengeti National Park, Tanzania, East Africa, Africa

Three lesser flamingo (Phoeniconaias minor) feeding with greater flamingo in the background, Serengeti National Park, Tanzania, East Africa, Africa

Immature red-billed oxpecker (Buphagus erythrorhynchus) on an impala, Kruger National Park, South Africa, Africa

Bison (Bison bison) bull demonstrating the flehmen response, Yellowstone National Park, Wyoming, United States of America, North America

Yellow pine chipmunk (Eutamias amoenus) eating a grass head, Yellowstone National Park, Wyoming, United States of America, North America

African elephant (Loxodonta africana) adult supporting a baby drinking, Addo Elephant National Park, South Africa, Africa

Cape mountain zebra (Equus zebra zebra) sparring, Mountain Zebra National Park, South Africa, Africa