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Greater kudu (Tragelaphus strepsiceros) buck with his mouth open, Kgalagadi Transfrontier Park encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Diamond Mine in extreme north-west, located on the southern bank of the Orange River mouth, Alexander Bay, Northern Cape, South Africa, Africa

Diamond Mine in extreme north-west, located on the southern bank of the Orange River mouth, Alexander Bay, Northern Cape, South Africa, Africa

Great Egret (Ardea alba) on left and little egret (Egretta garzetta) at Nosara Beach and river mouth, Nosara, Guanacaste, Costa Rica, Central America

Brown Pelican (Pelecanus occidentalis) looking for fish at Nosara Beach and river mouth, Nosara, Guanacaste Province, Costa Rica, Central America

Photographer Skip Brown on a stand up paddle board at the mouth of Havasu Creek in the Grand Canyon, Arizona, United States of America, North America

Monster Mouth Door, Structure II, Mayan Ruins, Hormiguero Archaeological Zone, Rio Bec Style, Campeche State, Mexico, North America

Monster Mouth Door, Structure II, Mayan Ruins, Hormiguero Archaeological Zone, Rio Bec Style, Campeche State, Mexico, North America

Monster Mouth Door, Structure II, Mayan Ruins, Chicanna Archaeological Zone, Campeche State, Mexico, North America

An evening view of the harbour and mouth of the River Teign, plus Shaldon Bridge, with Teignmouth on the left and Shaldon on the right, on the south coast of Devon, England, United Kingdom, Europe

Nile crocodile (Crocodylus niloticus) mouth open in the river Chobe, Chobe National Park, Botswana, Africa

An aerial view of the historic 16th century Dartmouth Castle, in the mouth of the River Dart, with a view of Dartmouth in the background, on the south coast of Devon, England, United Kingdom, Europe

Aerial view of beach and dunes at Dawlish Warren, guarding the mouth of the River Exe, looking south along the coast towards the town of Dawlish, Devon, England, United Kingdom, Europe

An aerial view of the historic 16th century Dartmouth Castle, in the mouth of the River Dart, at Dartmouth, on the south coast of Devon, England, United Kingdom, Europe

An aerial view of the historic 16th century Dartmouth Castle, in the mouth of the River Dart, Dartmouth, on the south coast of Devon, England, United Kingdom, Europe

Pharaoh Ay Performs the Opening of the Mouth Ceremony on Tutankhamun, North Wall, Burial Chamber, Tomb of Tutankhamun (King Tut), KV62, Valley of the Kings, Ancient Thebes, UNESCO World Heritage Site, Luxor, Egypt, North Africa, Africa

Waterfall pouring down a rocky cliff, with blurred motion, resulting from a slow shutter speed; at Welcombe Mouth, Hartland, north Devon, England, United Kingdom, Europe

Eype mouth, Dorset coast path to Thornecombe Beacon, Jurassic Coast, UNESCO World Heritage Site, near Bridport, Dorset, England, United Kingdom, Europe

A dusk view of historic buildings around the old harbour at Dartmouth, in the mouth of the River Dart, on the south coast of Devon, England, United Kingdom, Europe

A dusk view of Kingswear, seen from Dartmouth, in the mouth of the River Dart, on the south coast of Devon, England, United Kingdom, Europe

Waterfall pouring down a rocky cliff, with blurred motion, resulting from a slow shutter speed; at Welcombe Mouth, Hartland, north Devon, England, United Kingdom, Europe

Folded sedimentary rocks in cliffs at Welcombe Mouth, on the Atlantic coast; Hartland, north Devon, England, United Kingdom, Europe

Waterfall pouring down a rocky cliff, with blurred motion, resulting from a slow shutter speed; at Welcombe Mouth, Hartland, north Devon, England, United Kingdom, Europe

An aerial view of sand banks at low tide, in the mouth of the River Avon, at Bigbury, on the south coast of Devon, England, United Kingdom, Europe

South American black vulture (Coragyps atratus brasiliensis), a common scavenger, river mouth near Nosara, Nicoya Peninsula, Guanacaste Province, Costa Rica, Central America

The dramatic mixed sand and rocks of the beach at Speke's Mill Mouth, on Devon's Atlantic coast, near Hartland Quay, Devon, England, United Kingdom, Europe

Lion (Panthera leo) demonstrating the flehmen response, Ngorongoro Conservation Area, UNESCO World Heritage Site, Serengeti, Tanzania, East Africa, Africa

Fur seals sunning themselves on a rock at the mouth of Doubtful Sound, Fiordland, South Island, New Zealand, Pacific

Wall painting of the God Anubis performing the Opening the Mouth Ritual on Pharaoh, Tomb of Nebenmaat, TT219, Deir el-Medina Tombs (Worker Tombs), Thebes, UNESCO World Heritage Site, Luxor, Egypt, North Africa, Africa

Waterfall shot with a slow shutter speed, resulting in blurred motion, at Welcombe Mouth, Hartland, north Devon, England, United Kingdom, Europe

This fish is having its gills cleaned by a cleaner wrasse so it opens wide allowing the fish to come in and feed on parasites, Raja Ampat, Indonesia.

Guadalupe fur seals (Arctocephalus townsendi), at new haul out on Las Animas Island, Baja California Sur, Sea of Cortez, Mexico, North America

A rocky shore at low tide and under grey skies; at Welcombe Mouth, Hartland, north Devon, England, United Kingdom, Europe

Monumental statue of Buddha at Bentota River mouth, Bentota, Galle District, Southern Province, Sri Lanka, Indian subcontinent, South Asia

Woman holding a young alligator. Boat trip to explore the mangrove at the Bentota River mouth, Bentota, Galle District, Southern Province, Sri Lanka, Indian subcontinent, South Asia

A waterfall cascades down a gorge cut into coastal cliffs, at Speke's Mill Mouth, near Hartland Quay, on Devon's Atlantic coast, Devon, England, United Kingdom, Europe

Monumental statue of Buddha at Bentota River mouth, Bentota, Galle District, Southern Province, Sri Lanka, Indian subcontinent, South Asia

Structure II, monster mouth temple at sunset, Central Plaza, Chicanna Archaeological Site, Campeche State, Mexico

Aerial view of beach and dunes at Dawlish Warren, guarding the mouth of the River Exe, looking south along the coast towards the town of Dawlish, Devon, England, United Kingdom, Europe

Mouth of Truth (Bocca della Verita), Basilica of St. Maria in Cosmedin, Rome, UNESCO World Heritage Site, Lazio, Italy, Europe

Parks and boats line the mouth of the Thiou River at Lake Annecy, Annecy, Haute-Savoie, France, Europe

Aerial view of the mouth of the River Exe, seen from above Dawlish Warren, looking towards the town of Exmouth, Devon, England, United Kingdom, Europe

An aerial view of a sand bar at the mouth of the River Teign, Teignmouth, Devon, England, United Kingdom, Europe

Aerial view of beach and dunes at Dawlish Warren, guarding the mouth of the River Exe, looking south along the coast towards the town of Dawlish, Devon, England, United Kingdom, Europe

View of couple on trail at Storms River Mouth, Tsitsikamma National Park, Garden Route National Park, South Africa, Africa

Aerial view of beach and dunes at Dawlish Warren, guarding the mouth of the River Exe, looking south along the coast towards the town of Dawlish, Devon, England, United Kingdom, Europe

Greater kudu (Tragelaphus strepsiceros) buck with his mouth open, Kgalagadi Transfrontier Park encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

The rocky coast of Penlee Point, looking towards Rame Head and at the mouth of Plymouth Sound, east Cornwall, England, United Kingdom, Europe

Aerial view of the mouth of the River Exe, seen from above Dawlish Warren and looking towards the town of Exmouth, Devon, England, United Kingdom, Europe

A lioness (Panthera leo) moving a young cub by carrying it in her mouth, Masai Mara National Park, Kenya, East Africa, Africa

An aerial view of the historic 16th century Dartmouth Castle, in the mouth of the River Dart, with a view of Dartmouth in the background, on the south coast of Devon, England, United Kingdom, Europe

An evening view across the harbour and sandbar in the mouth of the River Teign, looking towards the Ness headland, Teignmouth, on the south coast of Devon, England, United Kingdom, Europe

Aerial view of beach and dunes at Dawlish Warren, guarding the mouth of the River Exe, looking south along the coast towards the town of Dawlish, Devon, England, United Kingdom, Europe

The Vizcaya Bridge, UNESCO World Heritage Site, a transporter bridge linking the towns of Portugalete and Las Arenas in the Biscay province, crossing the mouth of the Nervion River, Spain, Europe

An aerial view of the town and harbour of Teignmouth, sitting in the mouth of the River Teign, south coast of Devon, England, United Kingdom, Europe

Aerial view of the mouth of the River Exe, seen from above Dawlish Warren and looking towards the town of Exmouth, Devon, England, United Kingdom, Europe

Bull moose (Alces alces), in the snow with open mouth, Grand Teton National Park, Wyoming, United States of America, North America

Various types of sonf, mouth freshener, in aluminium pots, some coated in silver, in sonf stall in market, Kolkata, West Bengal, India, Asia

Aerial view of beach and dunes at Dawlish Warren, guarding the mouth of the River Exe, looking south along the coast towards the town of Dawlish, Devon, England, United Kingdom, Europe

An aerial view into the mouth of the River Dart, with Kingswear nearest, towards the right, and Dartmouth in the distance, south coast of Devon, England, United Kingdom, Europe

Aerial view of beach and dunes at Dawlish Warren, guarding the mouth of the River Exe, looking south along the coast towards the town of Dawlish, Devon, England, United Kingdom, Europe

Mouth of Truth (Bocca della Verita), Basilica of St. Maria in Cosmedin, Rome, UNESCO World Heritage Site, Lazio, Italy, Europe

Lion (Panthera leo) yawning, Ngorongoro Conservation Area, UNESCO World Heritage Site, Serengeti, Tanzania, East Africa, Africa

Lioness (Panthera leo) drinking, Kgalagadi Transfrontier Park encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Lioness (Panthera leo), Kgalagadi Transfrontier Park encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Warm afternoon sun at mouth of the River Otter at Budleigh Salterton, Devon, England, United Kingdom, Europe

A view across the Ness headland to the mouth of the River Teign and the harbour and tourist resort of Teignmouth, on the south Devon coast, England, United Kingdom, Europe

View of Storms River Mouth, Tsitsikamma National Park, Garden Route National Park, South Africa, Africa

Eype mouth, Dorset coast path to Thornecombe Beacon, Jurassic Coast, UNESCO World Heritage Site, near Bridport, Dorset, England, United Kingdom, Europe

A yellow baboon (Papio hamadryas cynocephalus) walking with some food in its mouth, Tsavo, Kenya, East Africa, Africa

An aerial view of the mouth of the River Teign, looking across to the Ness headland and the village of Shaldon, near Teignmouth, south coast of Devon, England, United Kingdom, Europe

Aerial view of the mouth of the River Exe, seen from above Dawlish Warren and looking towards the town of Exmouth, Devon, England, United Kingdom, Europe

Aerial view of the mouth of the River Exe, seen from above Dawlish Warren and looking towards the town of Exmouth, Devon, England, United Kingdom, Europe

Aerial view of the mouth of the River Exe, seen from above Dawlish Warren and looking towards the town of Exmouth, Devon, England, United Kingdom, Europe

Aerial of La Gueule du Monstre (the Monster's Mouth) cave, Rurutu, Austral islands, French Polynesia, South Pacific, Pacific