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Great Horned Owl (Bubo virginianus) in captivity, Arizona Sonora Desert Museum, Tucson, Arizona, United States of America, North America

African red-eyed bulbul (black-fronted bulbul) (Pycnonotus nigricans), Kgalagadi Transfrontier Park, encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Crowned plover (crowned lapwing) (Vanellus coronatus), Serengeti National Park, Tanzania, East Africa, Africa

Lakeland Farm by Wastwater, early morning, Wasdale, Lake District National Park, Cumbria, England, United Kingdom, Europe

Gray Wolf (Canis lupus) running in the snow in captivity, near Bozeman, Montana, United States of America, North America

Mears Peak and the Sneffels Range in the winter, Uncompahgre National Forest, Colorado, United States of America, North America

Leopard (Panthera pardus) walking through dry grass with his tail up, Kruger National Park, South Africa, Africa

Bison (Bison bison) bull in the winter, Yellowstone National Park, UNESCO World Heritage Site, Wyoming, United States of America, North America

Red fox (Vulpes vulpes) in the snow, Grand Teton National Park, Wyoming, United States of America, North America

Cactus and sandstone formations at dawn, Valley Of Fire State Park, Nevada, United States of America, North America

Wilson Peak at dawn with a dusting of snow in the fall, Uncompahgre National Forest, Colorado, United States of America, North America

Salmon-coloured sandstone wall with evergreens, Grand Staircase-Escalante National Monument, Utah, United States of America, North America

Cloud over the Sneffels Range in the fall, Uncompahgre National Forest, Colorado, United States of America, North America

St. Peters and Piazza San Pietro in the early morning, Vatican City, UNESCO World Heritage Site, Rome, Lazio, Italy, Europe

A mosaic pavement by Miro and building decorated with Umbrellas, on Las Ramblas, Barcelona, Catalunya, Spain, Europe

Group of four grotesque chimneys on the roof of La Pedrera (Casa Mila), and roof window details on an apartment block designed by Antonio Gaudi, UNESCO World Heritage Site, Passeig de Gracia, Barcelona, Catalunya, Spain, Europe

She Wolf sculpture dating from the 5th century BC, Romulus and Remus probably added later, Capitolini Museum, Rome, Lazio, Italy, Europe

Storage alcoves in the changing room of the Roman Baths, dating from AD 138, Cilurnum (Chesters Roman Fort), Hadrian's Wall, UNESCO World Heritage Site, Chollerford, Northumbrian National Park, England, United Kingdom, Europe

The Royal Air Force Bomber Command Memorial, Green Park, Piccadilly, London, England, United Kingdom, Europe

Wild horse (Equus Caballus) mare and foal, Green River, Wyoming, United States of America, North America

Yellow aspen trees reflected in Priest Lake in the fall, San Juan National Forest, Colorado, United States of America, North America

Orange aspens in the fall, San Juan National Forest, Colorado, United States of America, North America

The clock tower and the long wall showing the serpentine design, The Ducal Palace (Palazzo Ducale) (Reggia di Colorno), Colorno, Emilia-Romagna, Italy

Sunset at Locust Point, North Rim, Grand Canyon National Park, UNESCO World Heritage Site, Arizona, United States of America, North America

Rocks in the badlands at sunrise, Bisti Wilderness, New Mexico, United States of America, North America

Cinnamon black bear (Ursus americanus) feeding, Waterton Lakes National Park, Alberta, Canada, North America

Fern Falls, Coeur d'Alene National Forest, Idaho Panhandle National Forests, Idaho, United States of America, North America

Nile crocodile (Crocodylus niloticus) swimming, Serengeti National Park, Tanzania, East Africa, Africa

Several yellow-billed oxpecker (Buphagus africanus) on a Masai giraffe (Giraffa camelopardalis tippelskirchi), Serengeti National Park, Tanzania, East Africa, Africa

Chapman's zebra (Plains Zebra) (Equus burchelli antiquorum) drinking, Kruger National Park, South Africa, Africa

Swallow-tailed bee-eater (Merops hirundineus), Kgalagadi Transfrontier Park, encompassing the former Kalahari Gemsbok National Park, South Africa, Africa

Coyote (Canis latrans), Yellowstone National Park, UNESCO World Heritage Site, Wyoming, United States of America, North America

Hoary marmot (Marmota caligata), Glacier National Park, Montana, United States of America, North America

Grizzly bear (Ursus arctos horribilis), Glacier National Park, Montana, United States of America, North America

Fern Falls, Coeur d'Alene National Forest, Idaho Panhandle National Forests, Idaho, United States of America, North America

Bison (Bison bison) cows sparring, Yellowstone National Park, Wyoming, United States of America, North America

Sunset over an Alpine tarn at Stoney Pass, San Juan National Forest, Colorado, United States of America, North America

Bison (Bison bison) bull, Yellowstone National Park, Wyoming, United States of America, North America

Orange and white sandstone layers with colorful clouds at sunrise, Valley Of Fire State Park, Nevada, United States of America, North America

Black bear (Ursus americanus) in the snow, Yellowstone National Park, Wyoming, United States of America, North America

Eroded boulders at the Egg Factory, Bisti Wilderness, New Mexico, United States of America, North America

Black bear (Ursus americanus) in the spring, Yellowstone National Park, Wyoming, United States of America, North America

Three black bear (Ursus americanus) cubs of the year or spring cubs, Yellowstone National Park, UNESCO World Heritage Site, Wyoming, United States of America, North America

The Clock seen from the Bell Tower, St. Marks, Square, Venice, UNESCO World Heritage Site, Veneto, Italy, Europe

Grizzly Bear (Ursus arctos horribilis), Yellowstone National Park, Wyoming, United States of America, North America

Yellowstone River near Calcite Springs, Yellowstone National Park, UNESCO Wortld Heritage Site, Wyoming, United States of America, North America

Shams-al Emarat Towers (Edifice of the Sun), Golestan Palace, UNESCO World Heritage Site, Tehran, Iran, Middle East

Young Masai giraffe (Giraffa camelopardalis tippelskirchi) feeding, Selous Game Reserve, Tanzania, East Africa, Africa

Water thickknee (water dikkop) (Burhinus vermiculatus), Selous Game Reserve, Tanzania, East Africa, Africa

Blue columbine (Colorado columbine) (Aquilegia coerulea), San Juan National Forest, Colorado, United States of America, North America

Young Masai giraffe (Giraffa camelopardalis tippelskirchi), Selous Game Reserve, Tanzania, East Africa, Africa

Field of Namaqualand daisy (Jakkalsblom) (Dimorphotheca sinuata), Namakwa, Namaqualand, South Africa, Africa

Ccommon zebra (plains zebra) (Burchell's zebra) (Equus burchelli), Selous Game Reserve, Tanzania, East Africa, Africa

Ruaha hornbill (Ruaha red-billed hornbill) (Tanzanian red-billed hornbill) (Tockus ruahae), Ruaha National Park, Tanzania, East Africa, Africa

Yellow baboon (Papio cynocephalus) mother and days-old infant, Ruaha National Park, Tanzania, East Africa, Africa

Green-backed heron (striated heron) (Butorides striatus), Kruger National Park, South Africa, Africa

D'Arnaud's barbet (Trachyphonus darnaudii), Ngorongoro Conservation Area, Tanzania, East Africa, Africa

Lion (Panthera leo) cub rubbing against its mother, Ngorongoro Crater, Tanzania, East Africa, Africa

Building reflections with women in brightly coloured dresses on sidewalk near Galeria Riga, Riga, Latvia, Europe