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Adult harlequin duck (Histrionicus histrionicus) in full breeding plumage in the calm waters of Lake Myvatn, Iceland

Adult Eurasian oystercatcher (Haematopus ostralegus ostralegus) in full breeding plumage at Isfjardardjup, Iceland

Rothschild's giraffe (Giraffa camelopardalis rothschildi), breeding dominant male, Woburn Safari Park, England, United Kingdom, Europe

Adult and juvenile king penguins (Aptenodytes patagonicus), at breeding colony at Salisbury Plain, South Georgia, Polar Regions

Adult and juvenile king penguins (Aptenodytes patagonicus), at breeding colony at Salisbury Plain, South Georgia, Polar Regions

Adult southern rockhopper penguin (Eudyptes chrysocome) at breeding colony on Saunders Island, Falkland Islands, South America

Magellanic penguin (Spheniscus magellanicus), a pair of breeding penguins on their nest, Patagonia, Chile, South America

A raft of common murres (Uria aalge) at breeding site on South Marble Island, Glacier Bay National Park, Alaska, United States of America, North America

A raft of common murres (Uria aalge) at breeding site on South Marble Island, Glacier Bay National Park, Alaska, United States of America, North America

A raft of common murres (Uria aalge) at breeding site on South Marble Island, Glacier Bay National Park, Alaska, United States of America, North America

King penguin (Aptenodytes patagonicus) breeding colony at Salisbury Plain, South Georgia, Polar Regions

King penguin (Aptenodytes patagonicus) breeding colony at Salisbury Plain, South Georgia, Polar Regions

Sunrise on king penguin (Aptenodytes patagonicus) breeding colony at Gold Harbor, South Georgia, Polar Regions

King penguin (Aptenodytes patagonicus) courtship display at breeding colony in Gold Harbor, South Georgia, Polar Regions

Adult King penguin (Aptenodytes patagonicus) breeding colony at Gold Harbor, South Georgia, Polar Regions

King penguin (Aptenodytes patagonicus) head detail at its breeding colony at Gold Harbor, South Georgia, Polar Regions

King penguin (Aptenodytes patagonicus) adults establishing partners at breeding colony in Gold Harbor, South Georgia, Polar Regions

King penguin (Aptenodytes patagonicus) adults establishing partners at breeding colony in Gold Harbor, South Georgia, Polar Regions

King penguin (Aptenodytes patagonicus) adult with chick at breeding colony in Gold Harbor, South Georgia, Polar Regions

Southern elephant seal (Mirounga leoninar) weaners at breeding beach in Gold Harbor, South Georgia, Polar Regions

Pre-dawn light on elephant seals (Mirounga leoninar), at breeding beach in Gold Harbor, South Georgia, Polar Regions

Southern elephant seals (Mirounga leoninar), at breeding beach in Gold Harbor, South Georgia, Polar Regions

Antarctic shag (Leucocarbo bransfieldensis) in breeding plumage, Barrientos Island, Antarctica, Polar Regions

Chinstrap penguin (Pygoscelis antarcticus), leaping from the sea at breeding colony on Barrientos Island, Antarctica, Polar Regions

Antarctic shag (Leucocarbo bransfieldensis) in breeding plumage, Barrientos Island, Antarctica, Polar Regions

Gentoo penguins (Pygoscelis papua), breeding colony on Weincke Island, Naumeyer Channel, Antarctica, Polar Regions

Tufted puffin (Fratercula cirrhata), breeding colony at South Marble Island, Glacier Bay National Park, Alaska, United States of America, North America

Landscape for Breeding and Training of Ceremonial Carriage Horses at Kladruby nad Labem, UNESCO World Heritage Site, Pardubice Region, Czech Republic, Europe

Adelie penguins (Pygoscelis adeliae), breeding colony in a snowstorm at Brown Bluff, Antarctic Sound, Antarctica, Polar Regions

Adelie penguins (Pygoscelis adeliae), on an ice floe at a breeding colony on Joinville Island, Antarctica, Polar Regions

Adelie penguins (Pygoscelis adeliae), breeding colony in a snowstorm at Brown Bluff, Antarctic Sound, Antarctica, Polar Regions

Adelie penguins (Pygoscelis adeliae), leaping near breeding colony, Heroina Island, Danger Island Group, Antarctica, Polar Regions

King penguin (Aptenodytes patagonicus), breeding colony at Salisbury Plain, South Georgia, South Atlantic, Polar Regions

Adult black-browed albatrosses (Thalassarche melanophris), at breeding colony on New Island, Falklands, South America

Adult black-browed albatrosses (Thalassarche melanophris), in flight at breeding colony on West Point Island, Falklands, South America

Black-browed albatross (Thalassarche melanophris), chicks at breeding colony on Saunders Island, Falklands, South America

Adult black-browed albatrosses (Thalassarche melanophris), at breeding colony on West Point Island, Falklands, South America

Black-browed albatross (Thalassarche melanophris), chicks at breeding colony on Saunders Island, Falklands, South America

Black-browed albatross (Thalassarche melanophris), chick at breeding colony on Saunders Island, Falklands, South America

Black-browed albatross (Thalassarche melanophris), chicks at breeding colony on Saunders Island, Falklands, South America

Breeding elegant terns (Thalasseus elegans) return to colony on Isla Rasa, Baja California Norte, Mexico, North America

Adult southern rockhopper penguins (Eudyptes chrysocome), at breeding colony on West Point Island, Falklands, South America

Adult and okum boy king penguins (Aptenodytes patagonicus) at breeding colony at Gold Harbor, South Georgia, Polar Regions

Adult king penguins (Aptenodytes patagonicus) amongst chicks at breeding colony at Gold Harbour, South Georgia Island, Polar Regions

King penguin (Aptenodytes patagonicus) chicks (okum boys) at breeding colony at Gold Harbour, South Georgia, Polar Regions

Adult king penguins (Aptenodytes patagonicus) amongst chicks at breeding colony at Gold Harbour, South Georgia Island, Polar Regions

A rare melanistic king penguin (Aptenodytes patagonicus), feeding its okum boy chick at Fortuna Bay, South Georgia Island, Polar Regions

Adelie penguin (Pygoscelis adeliae), at breeding colony at Brown Bluff on the eastern side of the Antarctic Peninsula, Antarctica, Polar Regions

Adelie penguin (Pygoscelis adeliae), feeding chick at breeding colony at Brown Bluff, Antarctic Peninsula, Antarctica, Polar Regions

Adult dovekies (Little Auk) (Alle alle) at their breeding site amongst scree slopes at Rubini Rock, Tikhaya Bay, Hooker Island, Franz Josef Land, Russia, Arctic Ocean, Eurasia

Adult black-legged kittiwake (Rissa tridactyla) in combat with a second kittiwake near Alexander Island, Franz Josef Land, Russia, Arctic Ocean, Eurasia

Adult black-legged kittiwake (Rissa tridactyla) in flight near Alexander Island in Franz Josef Land, Russia, Eurasia

Chinstrap penguin (Pygoscelis antarctica), at breeding colony at Half Moon Island, Antarctica, Southern Ocean, Polar Regions

Adult chinstrap penguin (Pygoscelis antarctica), carrying rock in its beak at breeding colony at Half Moon Island, Antarctica, Polar Regions

Adult black-legged kittiwakes (Rissa tridactyla) in flight in the Svalbard Archipelago, Barents Sea, Norway, Arctic, Europe

Brunnich's guillemot (Uria lomvia) breeding and nesting site at Cape Fanshawe in the Svalbard Archipelago, Norway, Arctic, Europe

Brunnich's guillemot (Uria lomvia) in flight near breeding and nesting site at Cape Fanshawe in Svalbard, Norway, Arctic, Europe

Brunnich's guillemot (Uria lomvia) breeding and nesting site at Cape Fanshawe in the Svalbard Archipelago, Norway, Arctic, Europe

Brunnich's guillemot (Uria lomvia) breeding and nesting site at Cape Fanshawe in the Svalbard Archipelago, Norway, Arctic, Europe

Brunnich's guillemot (Uria lomvia) breeding and nesting site at Cape Fanshawe in the Svalbard Archipelago, Norway, Arctic, Europe

Brunnich's guillemot (Uria lomvia) breeding and nesting site at Cape Fanshawe in the Svalbard Archipelago, Norway, Arctic, Europe

Adult great skua (Stercorarius skua) with recent kill in the Svalbard Archipelago, Norway, Arctic, Europe

Male Great frigatebird (Fregata minor) in breeding plumage with red gular pouch, on Genovesa (Tower) Island, Galapagos, UNESCO World Heritage Site, Ecuador, South America

Male Great frigatebird (Fregata minor) in breeding plumage with red gular pouch, on North Seymour Island, Galapagos Islands, UNESCO World Heritage Site, Ecaudor, South America

Male Great frigatebird (Fregata minor) in breeding plumage with red gular pouch, on North Seymour Island, Galapagos Islands, UNESCO World Heritage Site, Ecaudor, South America

An Atlantic Puffin (Fratercula arctica), on a cliff edge in Borgarfjaroarhofn, Iceland, Polar Regions

A breeding Shag (Gulosus aristotelis) by its nest on cliffs at Braes peninsula, a red listed bird, The Braes, Portree, Skye, Inner Hebrides, Scotland, United Kingdom, Europe

Adult gentoo penguin (Pygoscelis papua), leaping off ice shelf at Cuverville Island, Antarctica, Southern Ocean, Polar Regions

Antarctic fur seal pup (Arctocephalus gazella) on Prion Island in the Bay of Isles on South Georgia, Southern Ocean, Polar Regions

King penguins (Aptenodytes patagonicus) on the beach at breeding and nesting colony at Salisbury Plain in the Bay of Isles, South Georgia, Southern Ocean, Polar Regions

King penguins (Aptenodytes patagonicus) on the beach at breeding and nesting colony at Salisbury Plain in the Bay of Isles, South Georgia, Southern Ocean, Polar Regions

King penguin (Aptenodytes patagonicus) with introduced reindeer at breeding and nesting colony at St. Andrews Bay on South Georgia, Southern Ocean, Polar Regions

Sunrise on king penguin (Aptenodytes patagonicus) breeding and nesting colony at Salisbury Plain in the Bay of Isles, South Georgia, Southern Ocean, Polar Regions

King penguin (Aptenodytes patagonicus) breeding and nesting colony at Salisbury Plain in the Bay of Isles, South Georgia, Southern Ocean, Polar Regions

King penguin (Aptenodytes patagonicus) adult and chick at breeding and nesting colony at Salisbury Plain, South Georgia, Polar Regions

Adelie penguins (Pygoscelis adeliae), at breeding colony at Brown Bluff on the eastern side of the Antarctic Peninsula, Antarctica, Polar Regions

Adult black-browed albatross (Thalassarche melanophrys) in Elsehul on South Georgia, Southern Ocean, Polar Regions

Adult black-crowned night-heron (Nycticorax nycticorax falklandicus) on Carcass Island in the Falkland Islands, South America

First year Adelie penguin (Pygoscelis adeliae) chick at breeding colony at Brown Bluff, Antarctica, Polar Regions

Adelie penguins (Pygoscelis adeliae) hauled out onto the ice near Adelaide Island, Antarctica, Polar Regions