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Bull elk (Cervus canadensis) demonstrating the flehmen response, Jasper National Park, Alberta, Canada, North America

Two bull elk (Cervus canadensis) sparring during the rut, Jasper National Park, Alberta, Canada, North America

Hiking at Grey Glacier (Glaciar Grey), Torres del Paine National Park, Patagonia, Chile, South America

View down rainswept Milford Sound, mountains obscured by cloud, Milford Sound, Fiordland National Park, UNESCO World Heritage Site, Southland, South Island, New Zealand, Pacific

Aspen leaf turning red, orange, and yellow, Uncompahgre National Forest, Colorado, United States of America, North America

Afternoon thunderstorm, South Rim, Grand Canyon National Park, UNESCO World Heritage Site, Arizona, United States of America, North America

Everglades National Park, UNESCO World Heritage Site, Florida, United States of America, North America

Female Williamson's sapsucker (Sphyrapicus thyroideus) on a feeding tree, Yellowstone National Park, Wyoming, United States of America, North America

Bamburgh Castle and marram grass (ammophila arenaria) lit by golden evening light, Bamburgh, Northumberland, England, United Kingdom, Europe

Tourists standing at a cold lava stream after an eruption of Tolbachik volcano, Kamchatka, Russia, Eurasia

Reflections in the Museum of Modern Art Building, Manhattan, New York City, New York, United States of America, North America

Grey Glacier (Glaciar Grey) and Grey Lake (Lago Grey), Torres del Paine National Park, Patagonia, Chile, South America

Orbicular batfish (Platax orbicularis) close-up, Ras Mohammed National Park, off Sharm el-Sheikh, Sinai, Red Sea, Egypt, North Africa, Africa

Volcanic boulders on Navajo sandstone, Grand Staircase-Escalante National Monument, Utah, United States of America, North America

Ifandana rock, the place from which the Betsileo fell preferring death to Merina domination, Madagascar, Africa

Young Columbian ground squirrel (Citellus columbianus), Waterton Lakes National Park, Alberta, Canada, North America

Adult warbler finch (Certhidea olivacea), Santiago Island, Galapagos Islands, UNESCO World Heritge Site, Ecuador, South America

Sandhill crane (Grus canadensis), Yellowstone National Park, Wyoming, United States of America, North America

Grey-breasted spurfowl (gray-breasted spurfowl) (grey-breasted francoli) (gray-breasted francolin) (Francolinus rufopictus), Serengeti National Park, Tanzania, East Africa, Africa

Icebergs in Grey Lake (Lago Grey) with Grey Glacier (Glaciar Grey) behind, Torres del Paine National Park, Patagonia, Chile, South America

Rock formation, Uchisar, Goreme National Park, UNESCO World Heritage Site, Cappadocia, Anatolia, Turkey, Asia Minor, Eurasia

Derwent Water under a cold sunrise in October, Lake District National Park, Cumbria, England, United Kingdom, Europe

Young common waterbuck (Ellipsen waterbuck) (Kobus ellipsiprymnus ellipsiprymnus), Kruger National Park, South Africa, Africa

Stone statues at Tomb of Khai Dinh, Hue, UNESCO World Heritage Site, Vietnam, Indochina, Southeast Asia, Asia

Grey dawn sky above Cullernose Point on the Northumbrian coast, Northumberland, England, United Kingdom, Europe

Grey-breasted spurfowl (grey-breasted francolin) (Francolinus rufopictus), Serengeti National Park, Tanzania, East Africa, Africa

African elephant (Loxodonta africana) standing in a small stream, Serengeti National Park, Tanzania, East Africa, Africa

Black and white cuckoo (Oxylophus jacobinus), Serengeti National Park, Tanzania, East Africa, Africa

African elephant (Loxodonta africana) mother and baby, Serengeti National Park, Tanzania, East Africa, Africa

Lava gulls (leucophaeus fuliginosus), Cerro Brujo, San Cristobal Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

Loggerhead shrike (Lanius ludovicianus), Pawnee National Grassland, Colorado, United States of America, North America

Juvenile blesbok (Damaliscus pygargus phillipsi), Mountain Zebra National Park, South Africa, Africa

Tranquil grey morning at Convict Lake in the Eastern Sierras, California, United States of America, North America

Abandoned croft beneath a stormy sky in the township of Manish on the east coast of The Isle of Harris, Outer Hebrides, Scotland

Congresso Nacional (National Congress) designed by Oscar Niemeyer, Brasilia, UNESCO World Heritage Site, Brazil, South America

Phyllidia exquisita nudibranch, Indo-west Pacific waters, grows to 50mm, Philippines, Southeast Asia, Asia

Two teenaged male African Elephant (Loxodonta africana) playing, Kruger National Park, South Africa, Africa

Hiking at Grey Glacier (Glaciar Grey), Torres del Paine National Park, Patagonia, Chile, South America

Adult great skua (Stercorarius skua) with recent kill in the Svalbard Archipelago, Norway, Scandinavia, Europe

Bighorn sheep (Ovis canadensis) ram, Clear Creek County, Colorado, United States of America, North America

Abandoned croft beneath a stormy sky in the township of Manish on the east coast of The Isle of Harris, Outer Hebrides, Scotland

Yellow-dotted trevally (Carangoides fulvoguttatus) shoal, Ras Mohammed National Park, Red Sea, Egypt, North Africa, Africa

Famous houses in Ange de Guernisac Street with Viaduct in the background, Morlaix, Finistere, Brittany, France, Europe

View of Felsenkirche (church) and the coastal town of Luderitz with its colourful Germanic architecture, Namibia, Africa

Female Kirk's dik dik (Kirk's dikdik) (Madoqua kirkii), Serengeti National Park, Tanzania, East Africa, Africa

Grey langur at Ruvanvelisaya Dagoba, Anuradhapura, UNESCO World Heritage Site, North Central Province, Sri Lanka, Asia

Lava gull (leucophaeus fuliginosus), Cerro Brujo, San Cristobal Island, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America