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An adult neotropic cormorant (Nannopterum brasilianum), on New River near the Mesoamerican archaeological site of Lamanai, Belize

A weathered, moss-covered stone statue in a lush forest garden, surrounded by ferns and foliage near Sanzen-in Temple, Kyoto, Honshu, Japan

A serene fountain within the historic Enryaku-ji Temple complex, centre of Tendai Buddhism, on Mount Hiei, UNESCO, near Kyoto, Honshu, Japan

An adult neotropic cormorant (Nannopterum brasilianum), on New River near the Mesoamerican archaeological site of Lamanai, Belize

Southern right whale (Eubalaena australis) calf surfacing near Puerto Pyramides, Golfo Nuevo, Peninsula Valdez, Argentina

Traditional Shinto torii gate with guardian lion-dog statues surrounded by autumn foliage on Mount Hiei, near Kyoto, Honshu, Japan

West Indian manatee (Trichechus manatus), on the sand near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

West Indian manatee (Trichechus manatus), with snorkeler near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

West Indian manatee (Trichechus manatus), with snorkelers near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

West Indian manatee (Trichechus manatus), on the reef near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

West Indian manatee (Trichechus manatus), on the reef near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

West Indian manatee (Trichechus manatus), with snorkelers near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

Green sea turtle (Chelonia mydas), feeding on the sand near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

Green sea turtle (Chelonia mydas), feeding on the sand near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

Green sea turtle (Chelonia mydas), surfacing for air near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

Green sea turtle (Chelonia mydas), with snorkeler near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

Green sea turtle (Chelonia mydas), with snorkeler near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

Glacial iceberg detail from ice calved off the LeConte Glacier near Petersberg, Southeast Alaska, United States of America

Glacial iceberg detail from ice calved off the LeConte Glacier near Petersberg, Southeast Alaska, United States of America

Glacial iceberg detail from ice calved off the LeConte Glacier near Petersberg, Southeast Alaska, United States of America

Glacial iceberg detail from ice calved off the LeConte Glacier near Petersberg, Southeast Alaska, United States of America

Glacial iceberg detail from ice calved off the LeConte Glacier near Petersberg, Southeast Alaska, United States of America

Glacial iceberg detail from ice calved off the LeConte Glacier near Petersberg, Southeast Alaska, United States of America

Glacial iceberg detail from ice calved off the LeConte Glacier near Petersberg, Southeast Alaska, United States of America

Adult brown bear (Ursus arctos) shaking water from its fur near whale carcass in Glacier Bay National Park, Alaska, United States of America

Gathering of at least two resident pods of killer whales (Orcinus orca) encountered near sunset, Johnstone Strait, British Columbia, Canada

Gathering of at least two resident pods of killer whales (Orcinus orca) encountered near sunset, Johnstone Strait, British Columbia, Canada

Mated bald eagle (Haliaeetus leucocephalus) pair on nest near the Brooks River, Katmai National Park, Alaska, United States of America

Adult male spruce (Franklin's) grouse (Falcipennis canadensis) near Brooks River, Katmai National Park, Alaska, United States of America

Adult brown bears (Ursus arctos) courtship behaviour, Brooks River, Katmai National Park near Bristol Bay, Alaska, United States of America

A well-scarred old male polar bear (Ursus maritimus), on a fresh bearded seal kill near Monacobreen Glacier, Svalbard, Norway

Young curious female polar bear (Ursus maritimus), on fast ice near Hinlopen Strait in the Svalbard Archipelago, Norway

Mother polar bear (Ursus maritimus) with cub of the year feeding near a fin whale carcass in Svalbard, Norway

Young curious female polar bear (Ursus maritimus), on fast ice near Hinlopen Strait in the Svalbard Archipelago, Norway

Young curious female polar bear (Ursus maritimus), on fast ice near Hinlopen Strait in the Svalbard Archipelago, Norway

Young curious female polar bear (Ursus maritimus), on fast ice near Hinlopen Strait in the Svalbard Archipelago, Norway

Basalt boulders and cliffs, Johnson Canyon east of Bainbridge Steel Dam Lake near Ash Fork, Kaibab National Forest, Arizona, United States of America

Basalt formation in Johnson Canyon east of the Bainbridge Steel Dam Lake near Ash Fork, Kaibab National Forest, Arizona, United States of America

Basalt boulders and cliffs, Johnson Canyon east of Bainbridge Steel Dam Lake near Ash Fork, Kaibab National Forest, Arizona, United States of America

Dry waterfall cliffs, Johnson Canyon between Bainbridge Steel and Stone Dams near Ash Fork, Kaibab National Forest, Arizona, United States of America

Old water pipe, Johnson Canyon between Bainbridge Steel and Stone Dams near Ash Fork, Kaibab National Forest, Arizona, United States of America

Adult black guillemot (Cepphus grylle) in summer plumage near Vigur Island in Isafjardardjup Bay, Iceland

Dry waterfall cliffs, Johnson Canyon between Bainbridge Steel and Stone Dams near Ash Fork, Kaibab National Forest, Arizona, United States of America

Panorama of Kaibab Lake from its south side, Kaibab National Forest near Williams, Arizona, United States of America, North America

Grotta della Poesia (Poetry Cave) natural pool among karsk formations, Roca archaeological site, near Melendugno, Puglia, Italy, Europe

Posing for photos with mountains and Lake Wakatipu near Queenstown, Otago, South Island, New Zealand, Pacific

Loch an Eilein, near Aviemore, Cairngorms National Park, Highlands, Scotland, United Kingdom, Europe

The River Tamar at Cotehele Quay, the border between Devon and Cornwall, near Gunnislake, Cornwall, England, United Kingdom, Europe

Mother and calf polar bear (Ursus maritimus) on first year sea ice in Olga Strait, near Edgeoya, Svalbard, Arctic, Norway, Scandinavia, Europe

Waves washing rock formation near coast of Porto Moniz and distant view of lighthouse on Ilheu Mole Island, Madeira, Portugal, Atlantic, Europe

Kicker Rock, a volcanic formation near the island of San Cristobal, a popular spot for snorkelling, Galapagos Islands, UNESCO World Heritage Site, Ecuador, South America

View of contrasting city skyline and Japanese Red Pine trees near the Imperial Palace of Tokyo, Tokyo, Honshu, Japan

Young woman by the torrent, Riva Valley (Val di Riva) (Reintal), near Ahrntal (Valle Aurina), South Tyrol (Alto Adige), Italy

Camels near Bedouin camp in Sharqiya Sands, formerly Wahiba Sands, desert region, Sultanate of Oman, Arabian Peninsula

Southern right whale (Eubalaena australis) adult surfacing near Puerto Pyramides, Peninsula Valdez, Argentina

Green sea turtle (Chelonia mydas), feeding on the sand near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

Green sea turtle (Chelonia mydas), feeding on the sand near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

Green sea turtle (Chelonia mydas), surfacing for air near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

Green sea turtle (Chelonia mydas), surfacing for air near Caye Caulker, inside the Mesoamerican Barrier Reef, Belize

Gathering of at least two resident pods of killer whales (Orcinus orca) encountered near sunset, Johnstone Strait, British Columbia, Canada

Float planes used to fly guests in to the Brooks Lodge and campground in Katmai National Park near Bristol Bay, Alaska, United States of America

An adult ivory gull (Pagophila eburnea) on ice near Monaco Glacier on the north side of SpitsBergen, Vestland, Norway

Dry waterfall cliffs, Johnson Canyon between Bainbridge Steel and Stone Dams near Ash Fork, Kaibab National Forest, Arizona, United States of America

Basalt boulders and cliffs in a section of Johnson Canyon halfway between the Bainbridge Steel Dam and Stone Dam near Ash Fork Arizona in the Kaibab National Forest.

Adult humpback whale (Megaptera novaeangliae), surfacing near researchers in Zodiac at sunset, Iceland

Adult Andean fox (Lycalopex culpaeus) near its den in the Andean Central Volcanic Zone, Antofagasta Region, Chile, South America