Female mantled howler monkey (Alouatta palliata) eating tree leaves in a north Pacific coast forest, Esperanza, Nosara, Guanacaste, Costa Rica

alouatta palliata foraging tree leaves tropical forest costa rica wildlife primate flora fauna eating herbivore nature animal behavior green biodiversity rainforest conservation habitat ecological balance ecosystem mammal climbing nutrition wild animals feeding habits jungle monkey lush vegetation natural environment biology study observation health canopy foliage species diet research interaction herbivorous